Monoclonal LO2
Overview of Monoclonal Antibodies Production
Focus on the production of monoclonal antibodies, using hybridoma cell lines as the primary method.
Definition of Monoclonal and Polyclonal Antibodies
Monoclonal antibodies: Identical antibodies produced from a single clone of B cells.
Polyclonal antibodies: Antibodies produced by multiple B cell clones in response to an antigen. They require immunization of an animal and are thus produced in a limited quantity.
Advantages of Monoclonal Antibodies
Unlimited Production: Hybridoma cell lines allow for immortal cell lines resulting in continuous production of monoclonal antibodies.
Custom Production: Example: Monash University provides a facility for custom monoclonal antibody production based on specific antigens.
Historical Context and Development
Historical Background:
Kola and Milstein invented the hybridoma technology in 1975.
Received a Nobel Prize within ten years for this invention.
The technique led to the discovery of various leukocyte surface antigens, some of which were previously unknown.
Key Contributor: Alan Williams, an Australian immunologist at Oxford, collaborated with Milstein to use the hybridoma technique to differentiate leukocytes.
Immunized mice with rat thymocyte membranes to provoke an immune response.
Led to the identification of antibodies such as CD4 in 1977.
Hybridoma Technology Process Overview
Basic Steps in producing monoclonal antibodies via hybridoma technology include:
Immunization of a mouse.
Fusion of spleen cells with myeloma cells to create hybridomas.
Selection and cloning of hybridomas producing desired antibodies.
Culture for unlimited antibody production.
Step-by-Step Hybridoma Production Process
Immunization
Immunize a mouse with antigen X.
The mouse mounts an immune response against the foreign antigen.
Isolation of Spleen Cells
Extract the spleen, a crucial lymphoid organ rich in plasma cells (antibody-producing cells).
Culture the spleen cells to obtain polyclonal antibodies (temporary as these cells are not immortal).
Fusion with Myeloma Cell Line
Characteristics of Myeloma Cells: Cancer cells that are immortal but do not secrete antibodies.
Myeloma cells lack an enzyme known as HGPRT (hypoxanthine-guanine phosphoribosyltransferase).
Fusion Process: Use a chemical agent to fuse spleen cells (immunized, antibody-secreting) with myeloma cells to create hybridomas.
Selection of Hybridomas
Culture cells in HAT media (Hypoxanthine-Aminopterin-Thymidine media).
Aminopterin selectively kills myeloma cells (due to lack of HGPRT), allowing only the hybridomas to thrive.
Spleen cells will die off after a short time because they are not immortal.
Hybridomas survive due to their immortal myeloma origin and may use the salvage pathway for purine synthesis due to the presence of HGPRT.
Cloning and Expansion
Isolate hybridoma cells into individual wells for cloning.
Allow the cells to grow and secrete antibodies needed for testing.
Antibody Selection
Use ELISA (Enzyme-Linked Immunosorbent Assay) to test for binding to antigen X.
Identify the optimal hybridoma clone producing the desired antibody (e.g., the first clone in our example).
Expansion of Selected Clone
Clone of B cell gives rise to monoclonal antibodies with identical specificity, isotype, and affinity.
Different isotypes have varying effector functions, crucial for selecting appropriate antibodies based on intended use.
Production Scales
Tissue Culture: Hybridomas can be cultured in research laboratories at small scales.
Bioreactors: For larger-scale production, hybridomas can be cultured in bioreactors to yield significant quantities of monoclonal antibodies suitable for clinical or commercial applications.
Conclusion on Hybridoma Technology
Hybridoma technology allows for the sustained, unlimited production of monoclonal antibodies, revolutionizing biomedical research and therapeutic developments.