M12 BIO
Module 12: Evolution and Speciation
Understanding Evolution
Evolution: Change in heritable traits over generations.
Sources of Variation:
Mutations: Random DNA changes (beneficial, neutral, harmful).
Sexual Reproduction: Shuffling genes.
Gene Flow: Alleles move between populations.
Genetic Drift: Random allele frequency changes (strong in small pops).
Key Ideas:
Evolution isn’t goal-directed.
Complexity ≠ evolutionary “better.”
Evidence for Evolution
DNA Analysis: Genetic similarities.
Fossils: Record of gradual changes.
Homologous Structures: Same structure, different function (e.g., limbs).
Vestigial Structures: Leftover traits (e.g., appendix).
Formation of New Species (Speciation)
Species: Group that interbreeds and produces fertile offspring.
Speciation: Formation of new species through genetic isolation + divergence.
Mechanisms of Speciation
Allopatric ("different homeland"):
Geographic separation.
Caused by dispersal or vicariance.
Sympatric ("same homeland"):
No physical barrier.
Often due to:
Polyploidy (extra chromosome sets) in plants.
Behavioral shifts or new ecological niches.
Adaptive Radiation
Rapid speciation to fill different niches (e.g., Hawaiian honeycreepers).
Reproductive Isolation
Prezygotic Barriers (before fertilization):
Temporal, habitat, behavioral, mechanical, gametic.
Postzygotic Barriers (after fertilization):
Hybrid inviability, hybrid sterility, hybrid breakdown.
Reconnection and Speciation Rates
Hybrid Zones:
Reinforcement, fusion, stability.
Speciation Models:
Gradualism: Slow, steady changes.
Punctuated Equilibrium: Rapid change, long stability.