Community Ecology Notes

Community Ecology Introduction

  • Community ecology explores biodiversity by examining species interactions and environmental factors.
  • A community is composed of interacting species within a specific space and time.
  • Community ecology studies the patterns of diversity, abundance, and composition of species.
  • Community ecologists formulate theories and test them by comparing predictions to data.

Measuring Biodiversity

  • Biodiversity refers to the variety of life at all levels of organization.
  • Metrics for characterizing communities include:
    • Species richness (number of species)
    • Abundance (number of individuals)
    • Evenness (homogeneity of abundances)
    • Diversity indices (e.g., Shannon's index H=<em>i=1Sp</em>ilnpiH = -\sum<em>{i=1}^{S} p</em>i \ln p_i)
    • Species composition (identity and relative abundance)
    • Turnover (changes in composition over space or time)

Fundamental Patterns of Biodiversity

  • Species-area relationships (SARs) describe the relationship between habitat area and the number of species.
    • S=cAzS = cA^z where:
      • SS = number of species
      • AA = area
      • zz = fitted exponent (slope of the curve)
      • cc = coefficient
    • More area generally correlates with more species.
  • Species abundance distributions (SADs) describe the abundance of each species in a community.
    • They can be represented graphically or as histograms.
    • SADs capture information about richness, evenness, and community structure.

Mark Vallon's Framework for Community Ecology

  • Organizes community ecology around four main processes:
    • Selection/niches: Species differences regarding resource use, environment responses, and interactions.
    • Dispersal: Movement of species through space.
    • Drift/demographic stochasticity: Randomness in birth and death events.
    • Speciation: Origination of new species.

Two Species Lotka-Volterra Competition Model

  • Describes the interaction between two species competing for the same resource
  • dN<em>1dt=r</em>1N<em>1K</em>1N<em>1α</em>12N<em>2K</em>1\frac{dN<em>1}{dt} = r</em>1N<em>1\frac{K</em>1 - N<em>1 - \alpha</em>{12}N<em>2}{K</em>1}
  • dN<em>2dt=r</em>2N<em>2K</em>2N<em>2α</em>21N<em>1K</em>2\frac{dN<em>2}{dt} = r</em>2N<em>2\frac{K</em>2 - N<em>2 - \alpha</em>{21}N<em>1}{K</em>2}
  • Where:
    • NiN_i = population density of species i
    • rir_i = per capita population growth rate of species i
    • KiK_i = carrying capacity of species i
    • αij\alpha_{ij} = competition coefficient, effect of species j on species i
  • Equilibria are found when dNidt=0\frac{dN_i}{dt} = 0
  • Four possible equilibria:
    • (0,0)(0,0) (both species extinct)
    • (K1,0)(K_1, 0) (species 1 dominant)
    • (0,K2)(0, K_2) (species 2 dominant)
    • (N<em>1^,N</em>2^)(\hat{N<em>1}, \hat{N</em>2}) (coexistence):N<em>1^=K</em>1α<em>12N</em>2\hat{N<em>1} = K</em>1 - \alpha<em>{12}N</em>2 , N<em>2^=K</em>2α<em>21N</em>1\hat{N<em>2} = K</em>2 - \alpha<em>{21}N</em>1
  • For coexistence equilibrium to be biologically sensible(positive abundances), either: \frac{K1}{K2} > \alpha{12} and K</em>2K<em>1>α</em>21\frac{K</em>2}{K<em>1} > \alpha</em>{21}, OR \frac{K1}{K2} < \alpha{12} and K</em>2K<em>1<α</em>21\frac{K</em>2}{K<em>1} < \alpha</em>{21}