Wk_2___1_Histology_of_the_Nervous_System

Histology of the Nervous System Neuroanatomy 1


Readings

  • The Human Brain: Chapter 1 (all)

  • Optional: Clinically Oriented Anatomy: Chapter 1, section on nervous system.

  • Histology Textbook Material: Any relevant content on the nervous system.


Objectives

  • Identify the cell types of the nervous system.

  • Understand the basic parts of a neuron and their functions.

  • Recognize the organelles found in neurons and their unique aspects.

  • Explain the axonal transport system: its components and importance.

  • Describe the major components of synapses.

  • Classify the major types of neurons and glia, their locations, and functions.


Review Question: Cell Types

  • What are the 2 basic types of cells of the nervous system?

    • Neurons – transmit electrical and chemical signals.

    • Neuroglia (glia) – supportive cells with various functions.


Neuronal Parts


Review Question: Neuronal Parts

  • What are the 3 basic parts of a neuron?

  1. Cell body: Central portion with organelles, metabolism and information processing.

  2. Axon: Long cylindrical structure that transmits information.

  3. Dendrites: Thin, tapering processes that receive information from other neurons.


Characteristics of the Cell Body

  • Also known as soma or perikaryon.

  • Shape: round/polygon.

  • Size: 5-100 μm.

  • Contents: nucleus, organelles, cytoplasm.

  • Function: Carries out metabolism, synthesis, and information processing.


Characteristics of Dendrites

  • Structure: Thin, branching processes, thicker proximally and tapering distally.

  • Contents: Most organelles found in dendrites except nucleus; abundant at proximal end.

  • Function: Receive information from other neurons; surrounded by synapses and glial cells.


More on Dendrites

  • Longitudinal Section: Features include Nissl bodies, mitochondria, and cytoskeleton for support.

  • Cross-section: Extensively branched with presence of microtubules and Nissl bodies.


Dendritic Spines

  • Small projections from the end of dendritic branches.

  • Function: Increase surface area for synapses; essential for memory and intellect.


Axons

  • Characteristics: Long cylindrical processes, typically one per neuron, less branching than dendrites.

  • Parts:

    1. Axon hillock: Site where the axon attaches to the cell body.

    2. Initial segment: Most excitable area of the axon.

    3. Collateral branches: Side branches of the axon.

    4. Terminal branches: End branches leading to synapses.

    5. Bouton: Expanded distal end of the terminal branch; forms synapse, releases neurotransmitters.


Characteristics of Axons

  • Longitudinal section: Includes mitochondria, microtubules, but no Nissl bodies.

  • Cross-section: Mainly mitochondria, cytoskeleton, and myelin, which acts as electrical insulation.


Organelles


Review Question: Organelles

  • What cellular organelles do we know?

  • Functions: Nucleus, Golgi apparatus, mitochondria, Nissl bodies, nucleolus providing insight into the neuron’s secretory roles.


##What are Nissl Bodies?

  • Stained ribosomes that appear blue/purple under Nissl dye.

  • High concentrations indicate active synthesis processes in neurons relevant for identification.


Review Question: Cytoskeleton

  • Components: Microtubules, neurofilaments, intermediate filaments.

  • Functions: Provide structural support, shape, and transportation within the neuron.


Function of Microtubules

  • Hollow cylinders made of tubulin; integrated with microtubule-associated proteins (MAPs).

  • Role: Act as "railroad tracks" for axonal transportation.


Neurofilaments

  • Intermediate filaments consisting of cytokeratin proteins for structural support within neurons.


Microfilaments in Neurons

  • Composed of actin protein.

  • **Functions:**Movement of substances to cell membrane, axonal growth, and molecule attachment.


Axonal Transport System

  • Essential for axonal survival; classified into fast (anterograde and retrograde) and slow types.

  • Fast transport moves organelles, vesicles, and growth factors at high speeds; poorly understood mechanisms.


Fast Axonal Transport

  • Involves motor proteins like kinesin (anterograde) and dynein (retrograde) that traverse microtubules.

  • Parts: Head (ATPase) and tail (cargo-attached).


Synapses

  • Definition: Connections between neurons or neurons and muscles/glands.

  • Most common type occurs between axon and dendrite.


Structural Components of Synapses

  1. Presynaptic Element: Typically axon releasing neurotransmitters.

  2. Postsynaptic Element: Receives neurotransmitters and responds variably.

  3. Synaptic Cleft: Gap between presynaptic and postsynaptic elements where neurotransmitters diffuse.


Characteristics of Neurons

  • Dendrites collect information, soma synthesizes macromolecules, and axons conduct information to other neurons.


Neuronal Classification

  • By Function: Sensory, motor, interneurons.

  • By Control: Somatic vs autonomic.

  • By Structure: Multipolar, bipolar, pseudounipolar, unipolar.


Types of Neurons by Shape

  1. Multipolar: Many dendrites; most common.

  2. Bipolar: Two processes; found in special senses.

  3. Pseudounipolar: One process; involved in somatosensory functions.

  4. Unipolar: Rare in humans; single process.


Types of Glia in PNS

  • Schwann Cells: Myelinate axons; support injury response and metabolic functions.

  • Satellite Cells: Surround peripheral neuron cell bodies; provide structural and metabolic support.


Types of Glia in CNS

  1. Oligodendrocytes: Myelination support and protection.

  2. Astrocytes: Regulation of ions and neurotransmitters; control blood flow and structural support.

  3. Microglia: Immune function, debris removal, respond to injury.

  4. Ependymal Cells: Line CNS ventricles, CSF circulation, and hormone transport.


Summary of CNS Components

  • Cell Types: Neurons, macroglia, microglia.

  • Functions and Locations: Specific roles in processing, support, and injury response within gray and white matter regions of the CNS.


References

  • Dalley AF, Agur AMR (2022). Clinically Oriented Anatomy, 9th Ed., Wolters Kluwer.

  • Vanderah TW, Gould DJ (2021). Nolte’s The Human Brain: An Introduction to its Functional Anatomy, 8th Ed. Elsevier.