Speciation
- Speciation: Process by which one species splits, creating new biological diversity.
- Microevolution: Changes in allele frequency within a population over time.
- Macroevolution: Broad evolutionary patterns above the species level, resulting from a series of speciation events.
- Speciation bridges micro- and macroevolution.
Biological Species Concept
- Species: A group of populations capable of interbreeding with viable, fertile offspring.
- Gene flow maintains genetic cohesion.
- New species arise when populations become reproductively incompatible.
- Reproductive isolation is key to new species formation.
Reproductive Isolation
- Reproductive isolation: Barriers preventing interbreeding between species.
- Hybrids: Offspring resulting from successful interbreeding between two species.
Prezygotic Barriers:
- Habitat isolation: Species in different environments.
- Temporal isolation: Different breeding times.
- Behavioral isolation: Differing mating rituals.
- Mechanical isolation: Physical incompatibility.
- Gametic isolation: Incompatible eggs and sperm.
Postzygotic Barriers:
- Reduced hybrid viability: Low survival rates.
- Reduced hybrid fertility: Sterility.
- Hybrid breakdown: Weak or sterile hybrid offspring.
Speciation Modes
- Allopatric speciation: Geographic separation leads to divergence.
- It involves mutation, natural selection, and genetic drift.
- Sympatric speciation: Occurs without geographic separation.
- Mechanisms include polyploidy, habitat differentiation, and sexual selection.
Polyploidy:
- More than two sets of chromosomes (common in plants).
- Autopolyploid: Multiple chromosome sets from one species.
- Allopolyploid: Hybridization of different species.
Habitat Differentiation:
- Use of different resources limiting gene flow.
Sexual Selection:
- Selection based on mate preference, drives reproductive isolation.
Hybrid Zones
- Areas where different species can meet and mate leading to:
- Reinforcement: Strengthening of reproductive barriers.
- Fusion: Weakening of reproductive barriers, merging species.
- Stability: Continued hybridization without full fusion.
Macroevolution
- Pattern of evolution above the species level over vast time scales.
Origin of Life
- Abiotic synthesis of small organic molecules.
- Joining of molecules forming macromolecules.
- Molecules packaged into protocells.
- Self-replication begins.
- Alkaline vents and meteorites introduces molecules.
Fossil Records
It provides a biased view favoring species that existed longer, were abundant/widespread, and had hard parts.
Geological Record
- Divided into eons (Hadean, Archaean, Proterozoic, Phanerozoic) and eras (Paleozoic, Mesozoic, Cenozoic).
Key Events:
- Oxygen revolution: Accumulation of atmospheric oxygen.
- Eukaryotes Originated 1.8 BYA by endosymbiosis
- Evolution of multicellular eukaryotes: Increasing diversity and complexity.
- Ediacaran period: Emergence of complex, soft-bodied organisms.
- Cambrian explosion: Sudden appearance of modern animal phyla.
- Colonization of land: Plants, fungi, and animals adapt to terrestrial environments.
Plate Tectonics
- Movement of plates causes habitat and climate change as well as allopatric speciation.
Mass Extinctions
- Events causing significant species loss.
Permian Extinction:
- Volcanic activity, global warming, ocean acidification.
- Cretaceous Extinction: Asteroid impact, habitat change, climate change.
Adaptive Radiation
- Rapid evolution of diverse species from a common ancestor.
- Often follows extinction events or the evolution of new traits.
Phylogeny
- Evolutionary History of a species or group represented by tree branching diagrams.
- Key concepts in tree building.
- Rooted tree: Includes a common ancestor.
- Branch point: Shows divergence of lineages.
- Sister taxa: Closest relatives.
- Basal taxon: Early diverging lineage.
- Polytomy: Unresolved lineage with multiple branches.
Taxonomy
- It’s a classification and naming of organisms which follows the Linnaean system.
- Hierarchical classification:
- Domain, Kingdom, Phylum, Class, Order, Family, Genus, Species.
- Binomial nomenclature (Genus species).
Phylogenetic Trees Interpretations
- They show decent not phenotype and branching pattern not timeline. They also don’t indicate that a species evolved from an adjacent taxon.
Phylogenies
They’re inferred from morphological and molecular data using homology versus analogy.
Cladistics:
- Uses shared homologous characteristics to infer phylogeny.
- Identify clades (monophyletic, paraphyletic, polyphyletic).
Characters:
- Shared derived (unique to a clade) vs. shared ancestral characters.