mitochondrial_disease flashcards

Page 1: Introduction

  • Date: 17 October 2024

  • Course: Applied Medical Sciences – Molecular Basis of Disease (MEDC0010)

  • Topic: Mitochondrial DNA diseases

    • Presenter: Jan-Willem Taanman

    • Institution: Department of Clinical and Movement Neurosciences, UCL Queen Square Institute of Neurology, University College London

Page 2: Overview of Mitochondrial DNA (mtDNA) Diseases

  • Definition:

    • Diseases caused by qualitative or quantitative abnormalities of mtDNA.

  • Key Areas Covered:

    1. Mitochondria: structure and function

    2. mtDNA: structure, function, replication, and genetics

    3. Defects of mtDNA and their association with disease

    4. Strategies to reduce transmission of mtDNA point mutations

Page 3: Mitochondria: Structure and Function

  • Mitochondria are highly dynamic organelles.

  • They undergo continual fission and fusion.

  • Visualization Techniques:

    • Live-cell stimulated emission depletion (STED) microscopy

    • Transmission electron microscopy

Page 4: Functions of Mitochondria

  • Essential Functions:

    1. Generation of ATP through oxidative phosphorylation.

    2. Expression of an integral genome.

Page 5: Structure of Mitochondrial DNA (mtDNA)

  • Characteristics of mtDNA:

    • Closed-circular, double-stranded DNA approximately 16.6 kb in length.

    • Somatic cell contains thousands of mtDNA copies.

  • Organization:

    • Arranged in nucleoprotein complexes known as nucleoids

    • Nucleoids dispersed throughout the mitochondrial network.

Page 6: Mitochondrial DNA (mtDNA) (Continued)

  • Features:

    • Encodes only 13 proteins.

    • All 13 proteins are essential subunits of oxidative phosphorylation enzymes.

    • Other mitochondrial proteins must be imported from the cytosol.

Page 7: Mitochondrial Proteins

  • Location of Proteins:

    • mtDNA-encoded proteins within the mitochondrion.

    • Nuclear DNA-encoded proteins synthesized in cytosolic ribosomes.

Page 8: Bioenergetic Pathways

  • Overview of the Oxidative Phosphorylation (OXPHOS) System:

    • Location: Cytosol, Matrix, Intermembrane space.

  • Components:

    • Involves various complexes (I to V) for ATP synthesis.

  • mtDNA and nuclear DNA (nDNA) roles in protein encoding:

    • Specific counts for each complex's proteins sourced from mtDNA and nDNA.

Page 9: Replication of mtDNA

  • Constant turnover of mtDNA molecules.

  • Replication happens consistently across cell cycle phases,

    • Continues in post-mitotic cells.

Page 10: Immuno-detection Techniques

  • Methods:

    • BrdU incorporation into replicating DNA for cell cycle independent replication.

Page 11: Immunocytochemical Detection

  • BrdU incorporation shows mtDNA replication independence from the cell cycle.

Page 12: Structure of mtDNA Strands

  • Distinctions in mtDNA duplex strands:

    • Heavy (H) and Light (L) strands with unique densities.

Page 13: D-loop Structure

  • Some mtDNA molecules form a three-stranded structure (D-loop),

    • Important control site for mtDNA replication and transcription.

Page 14: Initiation Complexes of mtDNA Replication

  • Components involved:

    • TFAM, POLRMT, TFB2M, other helicases, and more, each plays a role in the replication process.

Page 15: Patterns of Inheritance

  • Maternal inheritance of mtDNA is established through outcomes post-fertilization.

Page 16: Mechanisms of Maternal Inheritance

  • Key facts:

    1. High mtDNA copy number in egg cells (150,000–700,000) vs. low in sperm (<10).

    2. Sperm mtDNA is degraded shortly after fertilization.

Page 17: Summarization of mtDNA Defects

  • Diseases associated with mtDNA defects cover various mechanisms including mutations.

    • Focus on nuclear DNA mutations that affect mtDNA.

Page 18: Strategies to Reduce mtDNA Mutations

  • Overview:

    • Techniques include egg donation, prenatal diagnosis, and preimplantation genetic diagnosis.

Page 19: Ethical Considerations

  • Discussion surrounding implications of techniques to mitigate mtDNA mutations.

Page 20: Current Research and Future Directions

  • Continued research focuses on innovative strategies for managing maternal inheritance of mtDNA.

Page 21: Recap of Key Points

  • Established connections between mtDNA mutations and various diseases essential for the understanding of genetic disorders.