BIOL130 Lecture 11

Page 1: Introduction

  • Course: BIOL130 Human Physiology

  • Topic: The Urinary/Renal System focusing on the basic renal process.

  • Instructor: Giulia Gurun M.S. Ph.D. MCD Biology, UC Santa Cruz

Page 2: Understanding the Basic Renal Process

  • Multiple choice question regarding the renal process. Correct choice is:

    • E) Fluid is filtered from glomerular capillaries into Bowman's space.

  • Other options represent misconceptions about renal processes.

Page 3: Filtration in Nephrons

  • Nephrons filter small amounts of plasma:

    • Plasma moves from glomerular capillaries into Bowman’s space (filtrate).

    • Tubular Reabsorption: Substances removed from the filtrate back to the bloodstream.

    • Tubular Secretion: Substances added from blood to the filtrate.

  • Water reabsorption occurs, and excess is excreted in urine.

Page 4: Importance of Water Reabsorption

  • Discussion prompt on why most filtered water is reabsorbed.

    • Key reason: To maintain fluid balance and prevent dehydration.

Page 5: Plasma Filtration in Nephrons

  • Only ~20% of plasma entering glomerular capillaries is filtered into Bowman’s space.

Page 6: Importance of Limited Plasma Filtration

  • Discussion prompt on the significance of filtering only a fraction of plasma volume.

    • Key reason: To control composition and volume of plasma effectively.

Page 7: Types of Nephrons

  • Two main types of nephrons:

    • Cortical Nephrons:

      • Short loops of Henle and mainly peritubular capillaries.

      • Most common type.

    • Juxtamedullary Nephrons:

      • Long loops of Henle and vasa recta.

      • Involved in urine concentration, found between cortex and medulla (about 15% of nephrons).

Page 8: Capillaries Associated with Nephrons

  • Nephrons relate to:

    • Glomerular Capillaries: Specialized for filtration (high blood pressure).

    • Peritubular Capillaries: Allow exchanges for reabsorption and secretion.

Page 9: The Glomerulus

  • Glomerulus contains fenestrated capillaries:

    • Allows large amounts of solute-rich fluid to pass through.

    • Fluid is usually low in protein content due to size constraints.

Page 10: Filtration Membrane

  • Composed of podocytes with foot processes:

    • Generates filtration slits, working with fenestrae to filter blood.

    • Model for how fluid passes into Bowman’s space.

Page 11: Glomerular Filtration

  • Passive process driven by hydrostatic pressure:

    • High efficiency due to large surface area and high glomerular hydrostatic pressure.

    • Glomerular Filtration Rate (GFR): Volume filtered per unit time (~180L/day).

Page 12: Impact of High Blood Protein on GFR

  • Discussion on how conditions like hepatitis C leading to hyperproteinemia affect GFR.

    • Increased protein levels in blood could reduce filtration efficiency due to increased oncotic pressure.

Page 13: GFR Regulation

  • GFR is influenced by vascular changes in the afferent and efferent arterioles.

  • Physiological regulation adjusts net glomerular filtration pressure to maintain average GFR (180L/day).

Page 14: Calculating Filtered Load

  • Filtered load = GFR x plasma concentration of substance:

    • Example: Glucose at 1g/L results in 180g/day.

    • Comparison shows absorption vs secretion of substances.

Page 15: Break

Page 16: Regulation of Substance Transport

  • Transport of substances is regulated by:

    • Membrane channels/transporters in tubular epithelial cells.

    • Regulation occurs via hormones and local factors.

Page 17: Tubular Reabsorption

  • Movement from tubular lumen to blood:

    • Begins with the filtration entering the tubule.

    • Na+ reabsorption primarily occurs actively, influencing the reabsorption of other substances.

Page 18: Limits on Substrate Reabsorption

  • Transport maximum (Tm) limits the reabsorption rate of substances:

    • Hyperglycemia can lead to glucosuria when plasma glucose exceeds Tm.

Page 19: Tubular Secretion

  • Movement from blood (peritubular capillary) to tubular lumen:

    • Includes disposal of drugs, waste elimination, and pH control.

Page 20: Division of Labor in the Tubule

  • Main site of reabsorption is the proximal convoluted tubule:

    • Features microvilli and specific transport proteins for Na+ and other ions.

    • Distal convoluted tubule fine-tunes substance excretion.

Page 21: Understanding Water Reabsorption

  • Question regarding the nephron segment with the most water reabsorption:

    • Correct answer: A) The proximal tubule.

Page 22: Sodium and Water Renal Processes

  • Na+ and water are freely filtered:

    • Active transport of Na+ in all segments except descending limb.

    • Water follows Na+ reabsorption according to permeability.

Page 23: Coupling of Water and Sodium Reabsorption

  • Water follows as Na+, Cl-, and other ions are absorbed, occurring passively by osmosis.

Page 24: Water Movement Across Epithelium

  • Water reabsorption is determined by:

    • Presence of aquaporin water channels, highly expressed in proximal nephron.

    • Absence in collecting ducts without ADH activity.

Page 25: Muddiest Point

  • Students prompted to note unclear topics for further clarification during study or office hours.