Lecture Histology module 2 3rd year Urinary system 2024-2025 2

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  • Title: Urinary System

  • Confidentiality Note: For internal use only; unauthorized distribution prohibited.

  • Author: Laura CHINEZU, MD PhD, Histology Department

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  • Components of the Urinary System:

    • Kidneys:

      • Function:

        • Form and release/remove urine

        • Remove toxins from the bloodstream

        • Conserve salts, glucose, proteins, and water

        • Regulate blood pressure, hemodynamics, and acid-base balance

        • Endocrine function: produce renin, erythropoietin

        • Activate Vitamin D (1,25-(OH)2 vit D)

    • Ureters: Deliver urine to the bladder

    • Urinary Bladder: Stores urine

    • Urethra: Conveys urine out of the body

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  • Kidney Anatomy:

    • Shape: Bean-like

    • Dimensions: Approximately 10 cm by 6.5 cm by 3 cm

    • Associated Structures:

      • Upper pole: Adrenal gland

      • Capsule, Cortex, Medulla, Renal sinus, Ureter, Renal pelvis, Septa of Bertin, Renal artery, Renal vein, Papillae, Calyces

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  • Kidney Features:

    • Concave region: The sinus

    • Contents:

      • Fat

      • Hilum: where the kidney is penetrated by

        • Ureter

        • Renal vein

        • Renal artery

        • Lymph vessels

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  • Kidney Structure:

    • Capsule:

      • Outer layer of fibroblasts and collagen

      • Inner layer with myofibroblasts

    • Stroma: Interstitial tissue

    • Parenchyma:

      • Cortex

      • Medulla

      • Composed of nephron, collecting tubules, interstitium/stroma, blood vessels

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  • Myofibroblasts:

    • Contractility may aid in resisting pressure changes in kidney function

    • Capsule extends inward at hilum, forming connective tissue covering of sinus

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  • Kidney Regions:

    • Cortex:

      • Reddish brown (rich vascularization; 90-95% of blood vessels)

      • Granular due to glomeruli/renal corpuscles

    • Medulla:

      • Pale (5-10% of blood vessels)

      • Striated (longitudinal orientation of tubules)

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  • Detailed Structure: Various abbreviations and shorthand notations related to regions or components of the kidney (specifics may be cryptic without context).

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  • Histological Notations: Indicates structural features of cortex and medulla, with region numbers and other descriptors relevant for referencing.

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  • Renal Cortex:

    • Cortical columns (Bertin) separate renal pyramids.

    • Cortical labyrinth: Substance between two medullary rays.

    • Contains renal corpuscles and convoluted segments of proximal and distal tubules.

    • Associated blood vessels are also present.

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  • Renal Medulla:

    • Contains 6 to 12 renal pyramids (Malpighi).

    • Base oriented towards cortex (corticomedullary border) and apex towards hilum (renal papilla).

    • Medullary rays: longitudinal striations; includes Henle loops, renal tubes, and collecting tubules.

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  • Kidney Lobe Structure:

    • Minor calyx

    • Renal cortex

    • Renal medulla

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  • Hemi-section Diagram: Shows pyramidal shape, cortical components, medullary rays, and other anatomical features at the renal level.

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  • Renal Papilla:

    • Apex perforated by about 20 openings of the ducts of Bellini, called area cribrosa.

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  • Kidney Lobe Definition:

    • Consists of renal pyramid and associated cortical substance.

    • Medullary ray with nearby cortical labyrinth defines a kidney lobule.

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  • Renal Corpuscle:

    • Key components include the lobe, stellate vein, afferent arterioles, glomerulus, and surrounding vasculature (e.g., interlobular arteries/veins).

    • Arterioles give rise to peritubular capillary network and vasa recta.

    • Blood supply to cortex is terminal—no anastomoses.

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  • Renal Lobule Structure:

    • Medullary ray forms the axis of the lobule, comprising straight tubules and collecting duct connections.

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  • Nephron Types:

    • Subcapsular/cortical

    • Juxtamedullary

    • Intermediate

    • Nephron Function: Morphological and functional unit of kidney, composed of renal corpuscles and renal tubules.

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  • Renal Corpuscle Structure:

    • Composed of Bowman's capsule and glomerulus.

    • Vascular and urinary poles present, central role in nephron function.

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  • Glomerulus:

    • Tuft of fenestrated capillaries; afferent arterioles branch into this system, drained by efferent arterioles.

    • Represents an arterial capillary bed.

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  • Vascular Components:

    • Afferent/efferent arterioles, glomerular capillaries, peritubular and interlobular vessels.

    • Arterial structure supports kidney function and blood filtration.

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  • Blood Supply Pathway:

    • Renal artery branches into segmentary arteries leading to interlobar arteries and ultimately arcuate arteries, supplying the kidney structure.

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  • Blood Flow Mechanics: Follow the flow from cortical to juxtamedullary structures, influencing urine concentration along the nephron.

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  • Kidney Vasculature: Organization crucial for blood processing; includes named structures depending on location or shape.

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  • Venous Drainage:

    • Peritubular capillaries drain into interlobular veins and then into arcuate and interlobar veins.

    • Erythropoietin synthesis in the renal cortex influences red blood cell formation.

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  • Bowman's Capsule:

    • Composed of two epithelial layers: parietal (simple squamous) and visceral (podocytes).

    • Contains capsular space for ultrafiltrate.

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  • Renal Microanatomy:

    • Interactions between various renal tubular components are crucial for nephron function and blood filtration.

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  • Podocytes:

    • Form fenestrated epithelium around glomerular capillaries; have foot processes contributing to the filtration barrier.

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  • Filtration Slits:

    • Covered by filtration slit diaphragm, regulating glomerular fluid flux and maintaining filtration efficiency.

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  • Filtration Apparatus Components:

    • Nephrin: Key element of filtration slit diaphragm; supporting proteins influence podocyte integrity and function.

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  • Filtration Slit Hierarchy:

    • Physiological structures interrelate to maintain selective permeability during filtration.

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  • Glomerular Basement Membrane (GBM):

    • A thick common BM with specific properties regulating molecular passage based on size and charge.

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  • GBM Structure:

    • Consists of lamina rara externa, lamina densa (type IV collagen), and lamina rara interna, each contributing to filter function.

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  • Filtration Barrier Characteristics:

    • Outer lamina impedes negative charge passage; inner lamina reflects molecular similarity in composition to outer lamina.

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  • Endothelial Surface Layer:

    • Fenestrated endothelium of glomerular capillaries covered in a glycocalyx that aids in filtration and molecular sieving.

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  • Filtration Barrier Components:

    • The collaboration of endothelium, basement membrane, and podocytes creates the glomerular filtration barrier.

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  • Filtration Summary:

    • Blood filtration process occurs at the glomerulus, producing an ultrafiltrate for renal processing.

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  • Mesangium Location and Function:

    • Extracellular matrix and specialized mesangial cells support glomerular architecture, modulate distension, and partake in phagocytosis.

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  • Mesangial Cells Functions:

    • Support capillary networks

    • Regulate glomerular distension and inflammation response

    • Remove debris and maintain filtration integrity.

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  • Mesangial Matrix:

    • Mesangial cells modify the matrix and regulate glomerular blood flow and filtration.

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  • Renal Tubule Overview:

    • Proximal tubule divides into convoluted and straight regions, leading into the loop of Henle.

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  • Proximal Tubule Anatomy:

    • Composed of cuboidal epithelium adapted for ion and fluid exchange, characterized by a brush border and extensive basal striations.

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  • Reabsorption Statistics:

    • 65% of ultrafiltrate reabsorbed by proximal convoluted tubule involving specific proteins like Na/K-ATPase and AQP-1.

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  • Reabsorption Mechanisms:

    • Active sodium transport, passive water movement, amino acids, sugars reabsorbed by proximal tubule cells facilitated by microvilli and associated enzymes.

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  • Henle’s Loop Functionality:

    • Three regions with unique permeability characteristics allowing for concentration gradients in ultrafiltrate osmolality.

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  • Distal Tubule Structure:

    • Comprised of three regions, covered in cuboidal cells with fewer microvilli, involved in electrolytic exchanges.

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  • Distal Tubule Functions:

    • Active sodium and potassium transport and bicarbonate reabsorption with hormonal regulation (aldosterone).

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  • Countercurrent Exchange Mechanism:

    • Interaction between collecting ducts, Henle loops, and vasa recta concentrates urine.

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  • Transport Mechanisms Overview:

    • Summary of ion exchange and reabsorption processes from nephron to collecting ducts.

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  • Capillary Association in Nephrons:

    • Renal structures involve multi-layered cell types with complex functions related to filtration and absorption.

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  • Juxtaglomerular Apparatus:

    • Consists of macula densa, juxtaglomerular cells, and extraglomerular mesangial cells; critical for blood pressure regulation.

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  • Cell Types in Juxtaglomerular Apparatus:

    • Includes smooth muscle cells, macula densa, juxtaglomerular cells, and mesangial cells that regulate kidney function.

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  • Macula Densa Function:

    • Monitors NaCl levels and releases mediators affecting juxtaglomerular renin secretion and glomerular filtration rate (GFR).

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  • Juxtaglomerular Cells Structure:

    • Smooth muscle modifications located in afferent arterioles that secrete renin.

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  • Renin-Angiotensin-Aldosterone System Activation:

    • Triggered by factors such as low Na+ concentration, leading to the cascade of hormonal responses that manage blood pressure and sodium reabsorption.

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  • Pharmacological Agents:

    • Various drugs target the RAAS pathway for hypertension and other kidney-related issues.

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  • Extraglomerular Mesangial Cells:

    • Involved in signaling between macula densa and juxtaglomerular cells, regulating blood pressure and filtration processes.

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  • Functions of Juxtaglomerular Apparatus:

    • Acts as an endocrine sensor for blood volume, affecting renin release while monitoring tubular fluid composition.

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  • Filtration Monitoring:

    • Macula densa regulates GFR by controlling arteriole dilation and renin release based on sodium concentration feedback.

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  • Vascular Influences:

    • Renin secretion mechanism and various pathways linking renal hemodynamics with blood volume and electrolyte balance.

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  • ACE Role:

    • Enzyme essential in the conversion of angiotensin I to angiotensin II, influencing blood pressure regulation.

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  • Collecting Tubules Composition:

    • Connected to distal convoluted tubules and have roles in urine conveyance and modification.

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  • Collecting Duct Epithelial Composition:

    • Composed of principal and intercalated cells, each playing distinct roles in electrolyte and fluid balance.

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  • Principal Cells Function:

    • Involved in Na+, K+, and water transport, regulated by aldosterone and ADH; intercalated cells play roles in acid-base balance.

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  • Polycystin Proteins:

    • Critical for mechanosensation and function of principal cells in the collecting tubules; mutations correlate with ADPKD.

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  • Collecting Ducts:

    • Ducts of Bellini functional in urine drainage and connect further downstream structures within the renal context.

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  • Interstitial Kidney Tissue:

    • Supports renal structures and includes various cell types involved in extracellular matrix production.

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  • Excretory Passages:

    • Structures include minor/major calyces, renal pelvis, ureters, urinary bladder, and urethra.

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  • Urothelium:

    • Stratified epithelium present in excretory passages, impermeable while allowing distension.

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  • Transitional Epithelium Diagrams:

    • Illustrations of dome-shaped cells with urothelial plaques involved in bladder expansion and contraction.

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  • Dome-Shaped Cell Surface:

    • Represents structural adaptations for stretching and contraction during bladder filling.

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  • Ureter Structure:

    • Composed of three layers, allows urine transport from kidneys to bladder.

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  • Urinary Bladder Structure:

    • Similar layered structure to ureter; includes muscular coat allowing further distension and contraction.

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  • Urinary Bladder Features:

    • Summary of the bladder's structural relevance to urine storage and excretion.

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  • Publication Note:

    • International edition with marked restrictions on distribution and sale.