More Gram-Negative Rods
Bacteria 8: More Gram-Negative Rods
GNR Objectives
Outline and organize the following Gram-negative rods by specific criteria:
- Pseudomonas
- Burkholderia
- Stenotrophomonas
- Legionella
- FusobacteriumCriteria for organization include:
- Biochemical testing
- Morphology
- Natural environment
- Antibiotic resistance
- Associated diseases
GNR Vocabulary
Key terms related to Gram-negative rods:
- Sepsis
- Endotoxin
- Opportunistic
- Pseudomonas aeruginosa:
- Oxidase: positive
- Non-fermenter: Aerobic
- Exotoxin A: inhibits protein synthesis similar to diphtheria toxin
- Pathologies: nosocomial infections, disease in cystic fibrosis, hot tub folliculitis, urinary tract infections (UTIs), antibiotic resistance
- Other terms: Capsule, slime layer, biofilm
- Burkholderia:
- Burkholderia pseudo-mallei: causes melioidosis
- Stenotrophomonas maltophilia: associated with pneumonia; treated with trimethoprim-sulfamethoxazole
- Legionella: habitats include aquatic environments, associated with Legionnaires' disease and Pontiac fever
- Buffered charcoal-yeast extract (BCYE) agar: selective growth medium for Legionella
- Fusobacterium: anaerobic, causes oral cavity infections
Importance of Gram-Negative Rods
Significance:
- Particularly critical in patients with predisposing conditions (e.g., immunocompromised)
- Major contributors to sepsis and endotoxic shock
- Contains numerous highly antibiotic-resistant strains
- Generally classified as opportunistic pathogens
Pseudomonas aeruginosa
Characteristics:
- Gram-negative rod
- Oxidase-positive
- Non-fermentative; primarily aerobic but can switch to anaerobic metabolism
- Presence of LPS (lipopolysaccharides) endotoxin typical of other Gram-negative bacteria
- Exhibits Antiphagocytic properties via exopolysaccharide slime (alginate)
- Pigments: Often produces fluorescent pigments pyoverdin and pyocyaninEnzymes/Exotoxins:
- Extensive lytic enzymes including hemolysins, leukocidins, and proteases
- Exotoxin A: inhibits protein synthesis by ADP-ribosylating translation factor EF-2 (similar to diphtheria toxin)
- Exoenzyme S: another ADP-ribosyltransferase that transfers ADP-ribose from NADInfections:
- Nosocomial infections spread via contaminated equipment, surfaces, and respiratory apparatus
- Common in immunocompromised individuals, particularly those with cystic fibrosis and post-surgical or burn patients
- Pseudomonal bacteremia can result in skin lesions termed ecthyma gangrenosum
- Diagnosis: Culture; characteristic fruity odor and pigment production
- Treatment: Antibiotics; however, resistance is common in cystic fibrosis patients
Burkholderia
Characteristics:
- Gram-negative rods
- Oxidase-positive
- Non-lactose fermenters
- Motile and aerobic
- Found ubiquitously in environments like water and soilSpecies/Pathologies:
- Burkholderia cepacia: common cause of pulmonary infections in cystic fibrosis patients
- Burkholderia pseudomallei: endemic to SE Asia and parts of Central/South America, Middle East, Pacific, Africa; causes melioidosis
- Symptoms include cough, pleuritic chest pain, and skin infections; presents in acute pneumonia and potential septicemia
- Could function as a biowarfare agent
- Burkholderia mallei: causes glanders in horses, severe respiratory symptoms, and infection complicationsDiagnosis: Culture
Treatment: Variability in antibiotic effectiveness; resistance is common
Stenotrophomonas maltophilia
Characteristics:
- Formerly classified with Pseudomonas, now recognized separately
- Gram-negative rod, atmospheric omnipresence
- Distinguishing feature: production of two inducible metallo-β-lactamasesInfections:
- Primarily opportunistic—affecting immunocompromised individuals via lung infections and indwelling linesDiagnosis: Culture
Treatment: Trimethoprim-sulfamethoxazole; resistance encountered with aminoglycosides, penicillins, and third-generation cephalosporins
Legionella pneumophila
Characteristics:
- Gram-negative bacilli, motile, with both oxidase and catalase positive activity
- Contains over 25 species and 42 serogroups, with L. pneumophila serotype 1 responsible for most infections
- Aquatic saprophytes that thrive in amoebae and biofilmsTransmission: Infection occurs primarily through aerosolized water, such as from HVAC systems, cooling towers, and showers
Pathologies:
- Legionnaires' disease: severe pneumonia that can lead to multi-organ failure if untreated; characterized by non-productive cough, fever
- Pontiac fever: an epidemic flu-like condition with symptoms lasting several daysLPS: Lacks the endotoxic lipid A moiety typically found in Gram-negative bacteria
Diagnosis: Grown on Buffered Charcoal-Yeast Extract (BCYE) agar; direct immunofluorescence and specific antibody detection also employed
Treatment: Azithromycin for children and fluoroquinolones (e.g., levofloxacin, moxifloxacin) for adults
Prevention: Regular maintenance of air conditioning systems and application of biocidal compounds to manage outbreaks
Fusobacterium
Characteristics:
- Gram-negative fusiform bacilli, strictly anaerobic
- Typically non-lactose fermenting and colonizes oral cavity as commensals
- Has potent LPS endotoxin and produces various other toxinsPathologies:
- F. nucleatum: implicated in dental plaque and periodontal disease
- F. necrophorum: leads to pharyngitis, tonsillitis, and peritonsillar abscesses; can cause Lemierre syndrome, presenting with septic thrombophlebitis due to pharyngeal infectionsDiagnosis: Anaerobic cultures, PCR
Treatment: Several effective antibiotics, though resistant to erythromycin and some strains produce β-lactamase
Summary of Bacteria of Interest
Key Species:
- Pseudomonas aeruginosa
- Burkholderia (including cepacia and other species)
- Stenotrophomonas maltophilia
- Legionella pneumophila
- Fusobacterium
Pseudomonas aeruginosa Infectious Profile
Importance: Notable for widespread infections across body systems, including:
- Lungs in immunocompromised patients (particularly cystic fibrosis)
- Skin infections from burns and folliculitis from contaminated water sources
- UTIs and line-related infections leading to systemic sepsisEnvironmental Sources: Present in water, medical equipment, and patient care environments
Virulence Factors of Pseudomonas aeruginosa
Primary Characteristics:
- Gram-negative, aerobic, oxidase positive, non-lactose fermenting
- Notable virulence components:
- Capsule, slime biofilm
- Secretion of numerous lytic enzymes
- Exotoxin A: inhibits ribosomal function impacting protein synthesis
- LPS endotoxin contributes to various pathogenic processes
Treatment and Resistance Patterns
Treatment Challenges: Increasingly multidrug-resistant (MDR); common antibiotics include aminoglycosides and penems, with resistance often noted in cystic fibrosis patients
Laboratory Identification: Bacterial cultures yield distinct characteristics: a grape-like scent and pigment production, particularly the blue-green from pyocyanin
Questions and Case Studies
Clinical Cases: Utilized in assessing knowledge and application of information regarding Gram-negative rod infections and appropriate treatment strategies: 1. Sepsis from a non-lactose fermenting organism 2. Urinary infections with notable characteristics 3. Wound infection scenarios with methicillin-resistant organisms
Educational Focus: Explore resistance mechanisms, appropriate diagnostics, and treatment considerations across different presentations of Gram-negative infections, especially in critically ill or immunocompromised patients.