Plant Biology and Environmental Biology Review
Plant Biology
Reproduction (Life Cycles)
- All plants alternate generations, meaning offspring have a different chromosome count than parents.
- Fertilization creates the diploid generation (sporophyte).
- Meiosis creates the haploid generation (gametophyte).
- Offspring have a different form from their parents, even though they are the same species.
Bryophytes
- Diploid generation alternates with haploid generation.
- The sporophyte (2n) is parasitic on the gametophyte (1n).
- The gametophyte is the "dominant" generation (larger, more conspicuous).
Ferns
- Diploid generation alternates with haploid generation.
- Both generations are photosynthetic and free-living (not parasitic).
- The sporophyte is the dominant generation.
Flowering Plants (Angiosperms)
- Diploid generation alternates with haploid generation.
- The gametophyte consists of only a few hundred cells in the flower.
- It is totally dependent on the sporophyte for nutrition.
- It makes gametes, which fuse and form the 2n seed (the seed is the embryo that grows into the adult sporophyte).
- The sporophyte is dominant.
Summary of Alternation of Generations
| Plant Type | Sporophyte (2n) | Gametophyte (1n) |
|---|
| Mosses | Parasitic | Photosynthetic dominant |
| Ferns | Photosynthetic dominant | Photosynthetic |
| Flowering Plants | Photosynthetic dominant | Parasitic inside flower |
Pollination
- Flower structure matches the pollinator.
Wind-Pollinated Plants
- Produce small, drab flowers.
- Make lots of pollen.
- Examples: pine trees, grass.
Butterfly-Pollinated Plants
- Flowers tend to be red or blue.
- Have long, curved nectar tubes.
- Little scent.
Moth-Pollinated Plants
- Pale flowers.
- Produce a sweet scent.
- Long, curved nectar tube.
- Open at night/evening; many close during the daytime.
Bee-Pollinated Plants
- Bees are attracted to yellow & blue colored flowers.
- Infrared patterns.
- Sweet scent.
- Nectar is easily accessible (short nectar tubes).
Fly-Pollinated Plants
- Drab flowers.
- Stinky smell.
Beetle-Pollinated Plants
- Pale color.
- Fruity or spicy scent.
- Some provide edible petals instead of nectar.
Hummingbird-Pollinated Plants
- Large, red flowers.
- Sturdy, straight nectar tube.
- Little scent.
Bat-Pollinated Plants
- Strong stem.
- Pale flower.
- Strong sweet or fruity scent.
Summary of Pollination
| Pollinator | Flowers | Nectar Tube | Scent | Other |
|---|
| Wind | Small drab | None | None | Lots of pollen |
| Butterflies | Red or blue | Long curved | Little | Open 24 hours/day |
| Moths | Pale | Long curved | Strong sweet | Open at night |
| Bees | Yellow or blue | Short | Sweet | Infrared patterns |
| Flies | Small drab | Short | Stinky | --- |
| Beetles | Pale | Moderate | Fruity or spicy | --- |
| Hummingbirds | Large red | Long (straight) | Little | Bill shape matches flower |
| Bats | Large pale | Moderate | Strong sweet/fruity | Strong stems or hanging |
Hormones
Auxin
- Geotropism (Gravitropism):
- Bending of stem upwards – negative geotropism.
- Bending down of root downwards – positive geotropism.
- Phototropism:
- Growth toward light in stems – positive.
- Auxin is more concentrated on the dark side.
- Stimulates cell elongation.
- Apical Dominance:
- Stem tip produces auxin.
- Auxin travels down the stem.
- Inhibits side branches.
- Example: avocado vs. bushes.
Gibberellins
- Promote mitosis and starch metabolism.
- Too much causes the plant to grow to death (no starch reserves).
Ethylene
- Ripening (produced by damaged tissue).
- Produced by damaged tissue.
Plant Anatomy
Stem
- Apical meristems (buds) make new cells (mitosis).
- Pith (“ground cells”) for support.
- Vascular bundles:
- Are packets of tubes for fluid transport.
- Vascular cambium:
- In some plants.
- Makes more xylem & phloem.
- Xylem:
- Moves water and minerals up the stem.
- Tube cells are hollow cell walls.
- Water is a “chain” because of hydrogen bonds:
- Pulled by evaporation in the leaves.
- Chain would theoretically break at 450 feet.
- Tallest tree = coastal redwood named Hyperion: 379.1 feet.
- Phloem:
- Cells are alive when functional (unlike xylem).
- Moves molecules up AND down.
- Transports sap: sugars, hormones, etc.
- Mechanism unknown.
- Sap (sugars) is mobilized from roots in spring.
- Woody stems:
- No vascular bundles.
- Cylindrical layers of tissue.
- External bark made by cork cambium.
- Rings are produced by different sizes of spring & summer cells.
Roots
- Epidermis.
- Pith.
- Pericycle prevents water loss.
- Xylem, phloem, perhaps vascular cambium in the middle.
Leaves
- Cuticle:
- Upper epidermis: top "skin" layer.
- Mesophyll:
- Palisade: does most of the plant’s photosynthesis.
- Spongy:
- Provides ventilation.
- Air needed for its carbon dioxide.
- Most efficient way to:
- Moisten surface for dissolution of gasses.
- Allow evaporation from inside leaf (evapotranspiration).
- Lower epidermis:
- Lower “skin.”
- With stomates (pores, holes):
- Prevents excessive water loss (a necessary evil).
- Guard cells open & close stomate:
- Lots of water: guard cells swell & open hole.
- Little water: cells collapse over hole.
- Vessels:
- Xylem on top of a leaf vein.
- Phloem on bottom.
- No vascular cambium in leaf.
Photosynthesis
- Remember, plants respire too!
- CO2 +
ewline sunlight + H2O \rightarrow sugars + O_2
- Uses light
- Visible light: 380 to 750
- Action spectrum = what is used for photosynthesis – predominantly blues and reds
- Light dependent reactions
- Converts sunlight to bonds in ATP & NADPH
- H2O \rightarrow O2
- ADP & P -> ATP
- NADP -> NADPH
- light
- Energy is now “fixed”- also called fixation
- light-independent reactions
- Calvin cycle or dark reactions
- CO_2 \rightarrow sugar
- ATP -> ADP & P
- NADP H -> NADP
- Don't need light, just the products of the light reactions
- Convert energy in ATP and NADPH to a more stable form
- Photosynthesis summary
- H2O \rightarrow O2
- CO2 \rightarrow C6H{12}O6
- light energy
Diversity
Viruses
- Some biologists consider them to be alive.
- Not classified into classes, etc.; few detectable traits.
- Structure:
- Protein coat (capsule or capsid): round or hexagonal.
- Nucleic acid (DNA or RNA): linear, 1 piece.
- Life cycle:
- No metabolism or growth of its own.
- Parent virus injects its nucleic acid into host (=behavior).
- "Normal" viruses (DNA):
- Virus’ DNA is attached to hosts DNA.
- Host energy & enzymes make DNA copies & protein coats.
- Retroviruses (RNA):
- Inject their RNA.
- Host cell makes DNA based on this RNA (“reverse transcription”).
- New DNA incorporated into host DNA.
- Rest as in DNA viruses.
- Example: HIV, Coronavirus
Domain Bacteria (= D. Eubacteria)
- Are prokaryotes:
- A simple type of cell.
- No internal membranes.
- Structure:
- Cell wall.
- Cell membrane (plasma membrane).
- DNA (= chromosome; naked & circular).
- Ribosomes (small, simple).
- Flagellum in some: “9+0” in structure.
- Reproduction:
- Most cells are made by mitosis (asexual cell division).
- Sex: in some bacteria (“conjugation”)—meaning swap DNA pieces.
- Diversity:
- Classification is changing radically.
- Not sorted into kingdoms yet.
- “True bacteria”:
- About 26 different types aka divisions.
- Most of these are heterotrophic: non-photosynthetic (absorb molecules for energy).
- Important in decay, nutrient cycles, infections.
- Division Cyanobacteria:
- “Cyanophytes” or “blue-green-algae.”
- Photosynthetic.
- Some can make ammonium from N_2 in the air:
- N needed for plants: in fertilizer (nitrate).
- N \equiv N only can be broken by lightning, industry, cyanobacteria.
- N_2 "fixed" in heterocyst into a dissolved form
Domain Archaea (= Domain Archebacteria)
- Are prokaryotes.
- But are very different from Bacteria:
- Different photosynthetic pathway and pigment.
- Different phospholipid in membrane.
- rRNA of several types, one like eukaryotic rRNA.
- DNA is circular, but with histones.
- Found mainly in extreme environments.
- Diversity:
- Four divisions are thermophiles: found in hot waters (like geysers), important for rapid high-temperature DNA copying.
- Another division includes:
- halophiles: found in very salty waters (ex: Great Salt Lake).
- methanogens: found in oxygen-free muds (swamps).
Summary of Domains
| Character | D. Bacteria (prokaryotic) | D. Archaea (prokaryotic) | D. Eukarya (eukaryotic) |
|---|
| Cell size | 1-10 μm | 1-10 μm | 5-100 μm |
| Cell wall | yes | yes | some |
| Internal membranes | never | never | always |
| Chromosomes | single, circular | single, circular | multiple, linear |
| Histones | never | some | always |
| Chlorophyll | dissolved | dissolved | on membranes |
| Chloroplasts | never | never | some |
| Mitochondria | never | never | always |
| Flagella | 9+0 | variable | 9+2 |
| Ribosomes | small simple | variable | large complex |
Domain Eukarya (Eukaryota)
Kingdom Protista
- Are miscellaneous eukaryotes, unicells & colonies.
- "Algae"
- Life cycle
- Most cells produced asexually (mitosis)
- Sex has been observed in most species: genetic recombination (swapping of chromosomes)
- Alternation of generations
- Two-or three-phase
- Alternating generations often look different-but not always
- Diversity (27,000 species)
- Six divisions based on pigments and energy storage
- Division Chlorophyta (green algae) ex: Spirogyra, Chara
- Division Bacillariophyta (diatoms) ex: Aulacoseira, Navicula
- Importance
- Oxygen production: about 60% from oceans (particularly the algae from it)
- Food: autotrophic (self-feeding = photosynthetic)
- Base of aquatic food chains
- Human consumption: kelp, ice cream, Jello
- Medicines: iodine
- Other: agar, gels
- "Protozoa”
- Heterotrophic (non-photosynthetic), mostly unicells
- Reproduction: mitosis and sex (in most)
- Diversity (28,000 species)
- Eleven phyla based on propulsion and spores
- Phylum Ciliophora (cilates): Paramecium
- Phylum Rhizopoda (amoebas): Amoeba
- Importance
- Most are harmless or beneficial (ex: in termites)
- Diseases: malaria, yellow fever, cholera
- "Slime molds" (580 species)
- Two divisions
- Usually unicellular, heterotrophic
- Some are multinucleate
- Some species aggregate when the environment goes bad
- Ex: dry or cold
- Then produce stalks and spores
- Spores survive until conditions improve
Kingdom Fungi
- Once classified with K. Plantae, but
- Heterotrophic
- Cell walls with chitin, not cellulose
- Most with sex
- Some are symbiotic: “together living” of fungus and alga
- Fungus gets food (sugars from photosynthesis)
- Alga gets protection, water & nutrients
- These are lichens
- Diversity (82,000 species)
- Five divisions
- True molds, yeasts & morels, mushrooms, athlete's foot & Penicillium, aquatic molds
- Importance
- Decay
- Help most trees absorb water and nutrients
Kingdom Plantae
- General
- Virtually all are photosynthetic
- All alternate generations
- One haploid (gametophyte), one diploid (sporophyte)
- Always look different
- "Nonvascular plants"
- No true vessels
- Gametophyte is dominant
- One division: D. Bryophyta (16,000 species)
- Mosses, liverworts
- "Lower vascular plants"
- Low-growing
- Division Lycophyta (1200 species): club mosses
- Division Monilophyta (13,000 species): ferns, horsetails
- "Gymnosperms"
- Nonflowering vascular plants (tracheophytes)
- Diversity (900 species in 4 divisions)
- D. Ginkgophyta
- Only Ginkgo biloba
- Entire population was only 7 trees
- D. Coniferophyta (550 species): conifers
- Wind pollinated (makes lots of pollen)
- Life cycle similar to angiosperms, but no flowers
- Includes firs, pines, cedars, redwoods, cypress
- "Angiosperms"
- The flowering plants
- Division Anthophyta only (235,000 species)
- C. Monocotyledones (65,000 species): “monocots”
- Leaves with parallel veins
- Petals in threes
- Includes grass, palm, lily, orchid, etc.
- C. Dicotyledones (170,000 species): “dicots”
- Leaves with branching veins
- Petals in fours or fives
- Includes oaks, pecans, herbs, poison ivy, etc.
Kingdom Animalia
- General
- All are multicellular (except one species)
- Heterotrophic
- Do not alternate generations
- Phylum Porifera (sponges. 5000 spp.)
- Colonial
- No muscles or nerves
- Six cell types
- Phylum Cnidaria (9000 species)
- Colony of interdependent cells
- Specialized cells: muscles, nerves, stinging
- Locomotion
- Includes jellyfish, corals, anemones
- Phylum Nematoda (10,000 species)
- Roundworms
- Complete digestive system
- Protective cuticle
- Mostly predators, some parasites
- Phylum Platyhelminthes (13,000 species)
- Flatworms
- Flukes, tapeworms
- Phylum Annelida (12,000 species)
- Segmented worms
- Polychaetes, leeches, oligochaetes (earthworm)
- Other “worms” (500,000 species): 13 phyla, mostly deep sea
- Phylum Mollusca (47,000 species)
- Soft body, often with shell
- Mostly marine
- Snails, clams, cephalpods (octopus), etc.
- Phylum Rotifera (1800 species): “wheel animalicules”
- Phylum Arthropoda (850,000 species)
- Chitin exoskeleton, jointed legs
- C. Arachnida: spiders, mites, scorpions
- C. Crustacea: shrimp, crabs, pillbugs
- C. Merostomata (4 species) horseshoe crabs: Limulus polyphemus
- C. Insecta (750,000 species) insects: beetles, ants, wasps, dragonflies, butterflies…
- Phylum Echinodermata (6000 species)
- Radial symmetry (five directions)
- Initial development like chordates
- Urchins, starfish, sand dollars
- Phylum Chordata: chordates
- Rigid dorsal notochord or backbone
- “Lower chordates” (1300 species): barrel-like
- sP. Vertebrata (41,700 species)
- “Fish” (20,000 species in 3 classes): lampreys, sharks & rays, bony fish
- Class Amphibia (2500 species)
- Skin and eggs must be moist
- Frogs, salamanders
- Class Reptilia (6000 species)
- Scaly skin and shelled eggs resist drying
- Lizards, snakes, turtles, crocodiles
- Class Aves (8600 species)
- Feathers, warm-blooded
- Birds
- Class Mammalia (4500 species)
- Subclass Prototheria (3 species)
- Poor temperature regulation
- Egg layers
- Platypus, spiny anteaters
- Subclass Metatheria (260 species)
- Moderate temperature regulation
- Marsupials
- Opossum, kangaroos, wallabys, etc
- Subclass Eutheria (4240 species)
- Good temperature regulation
- “Higher mammals”
- Sixteen orders: bats, moles, lemurs, rabbits, carnivores, anteaters, rodents, deer & cows, horses, elephants, manatees, cetaceans, primates.
Environmental Biology
Atmospheric Issues
Urban Air Pollution
- From combustion.
- Mainly HC’s, CO_2
- General symptoms: headache, nausea, lung disease, cancer.
- Cleaning = prevention
- Particulates: filters.
- Chemicals: scrubbers, precipitators, incinerators.
Indoor Air
- Radon (carcinogen)
- Decays to radioactive materials.
- Common in granitic areas.
- Here, about the national average.
- Fireplaces
- CO, CO_2, particles of ash.
- Heavy metals: from wrapping paper, comics.
- Paint fumes
- Bathroom cleaners
- Much worse due to increased airtightness
Stratospheric Ozone
- Ozone absorbs UV radiation.
- Destroyed by CFCs from air conditioners, foam, etc.
- Only two-thirds of all CFCs have reached the upper atmosphere.
- Predictions (as of 2010)
- 900,000 more nonfatal cancers per year (especially skin cancer).
- 30,000 more fatal cancers per year.
- $2 billion damage to crops, plastics, etc.
- Antarctic ozone hole is closing - complete healing around 2065
Acidic Precipitation
- Components
- SO_x (sulfuric acid): from low-grade coals for power.
- NO_x (nitric acid): vehicles.
- Effects
- Weakens plants.
- In waters, kills fish, etc.
- In cities:
- Damages car finish.
- Dissolves limestone.
Climate Change (Global Warming)
- “Do you believe in climate change?” – better to ask, “Do you think that climate change is real?”
- Greenhouse gasses: mostly CO2, CH4
- Natural greenhouse effect: sunlight warms Earth, the atmosphere retains some heat.
- Humans: cars/vehicles in general (50%), industry (45%), fires (5%).
- CO_2 trends
- 1750: 210 ppm (parts per million).
- 2022 average: 417 ppm (highest in > 3 million years).
- Rising by 3 ppm/yr.
- Auto & oil industries now acknowledge their role
- Effects
- Temperature
- 2024: hottest year on record
- 2015-2024: hottest 9 years on record
- Since 1750, + 1.7 degrees C
- Sea level rise
- IPCC expects 2 ft before 2100: goal < 2 degrees C (3.6 degrees F)
- Will be complicated by ice sheet melting
- Alaskan glaciers: 12-inch rise
- Greenland glaciers: 20 feet rise of sea level
- Antarctica: 180 ft rise
- May not be gradual: if ice sheets slide into ocean
- Threatens cities: New York, Los Angeles, Miami, San Francisco, maybe oceanfront in Memphis after this
- Croplands: that feed about half the world’s population
- More & stronger hurricanes that will be longer and slower-moving.
- Shifting rainfall belt: desert in Missouri?
- Solutions
- Drive less, maybe more electric cars, but what source of electricity?
- Phase out coal, oil, natural gas in favor of solar, wind, nuclear
- Add a “greenhouse label” to products (like nutrition label information)
- Reduce deforestation and improve reforestation
Tropical Deforestation
- Rates
- At least 22,000 sq miles per year
- At least 2% of original rainforest area
- All of Missouri in 3 years
- Well over half gone now – probably 60%
- Reasons
- Huge debt of nation: sell permits to harvest wood
- Overpopulation: need cropland, but
- Crops fail after 3-4 years
- Pasture fails after 6 years
- Fuel gathering
- Results
- Massive erosion
- Enhanced greenhouse effect
- Loss of diversity
- 50-70% of all species in 7% of Earth's land area
- Potential food
- 90% of modern ones were originally tropical
- E.g., corn, potatoes, soybeans, tomatoes, peppers
- Potential medicines
- 40% of modern ones (digitalis, quinine)
- HIV, cancer cures?
- Global ecosystem stability
- Cultural diversity
- Solutions
- Slow population growth
- Forgive debt
- Trade debt for nature: environmental clubs, world bank, tropical rain forest country
- Enhance ecotourism
- Use mixed cropping (leave some trees intact)
Water Issues
Wastewater Pollution (Sewage)
- Is the biggest problem.
- Wastewater treatment plants: need upgrading.
- Feedlot sewage: poorly regulated.
Other Pollutants
- Industrial: organics, metals, heat.
- Land runoff: fertilizer, pesticides.
- Shipping: spills, bilgewater.
Water Use
- Use > resupply in Midwest wells
Solid Waste
Domestic
- 4.9 lbs/person/day (for 2019).
- Mainly paper (40%), garden (18%), food (12%), 8% each glass, wood, metals, plastics
Litter
- Why: it’s easy and free.
- Control
- More bins.
- Enforcement.
- Despite laws: reduces by half, saves energy
Negatives of Solid Waste
- Pests, disease, fires, blows away, cost.
Waste Reduction (3-R’s)
- Reduce: use less.
- Re-use: no reproduction of product (ex: deposit bottles).
- Re-cycling
- Re-production of the same product.
- Saves energy and cost of mining, refining.
- Cost is sorting & collecting.
- Paper: 1 ton = 17 trees
- Process: sort, liquify (melting or pulping), mold
- One product is made into a different one.
- E.g., plastics
- Re-formed into benches, insulation, etc.
- Can't re-form for food: cannot be sterilized.
- Sorted by code (7 types)
Disposal
- Composting: pile organics, let bacteria decompose.
- Incineration
- Generates heat, air pollution.
- Expensive
- Sanitary landfills
- Residue from all of the above, plus more.
- Covered each day.
- Problems: settling, rainwater, ugly, stink
Hazardous Materials
Types
- Radioactive: nuclear waste, hospitals, monitors.
- Heavy metals: lead, mercury in paints, industrial waste.
- Toxins (toxic waste)
- Industrial bleaches: stored in fat, released when dieting
- Pesticides
- Aim: complete eradication
- Problems
- Resistance: some pests survive, breed (natural selection)
- Biological amplification (magnification)
- Dangerous
- Long-lived
- Alternatives: aim is to minimize damage
- Release sterile mates
- Release predators
- Pheromones as attractant to traps
- Crop resistance
General Impacts
- Lesions, cancer, bones, kidney, nerves, liver.
Management of Hazardous Materials
- Cradle-to-grave tracking.
- Minimize use.
- Legal disposal methods
- High temperature
- Bury in super-safe landfill (24%)
- Deep well injection (14%)
- Theoretical disposal
- Shoot to sun
- Nursing in icecap
- 62% is disposed otherwise
- Superfund -- EPA
- 40,000 sites need work; 1,300 are urgent
- 428 in progress, 1031 completed
- Cost $400 to 500 billion
Energy Issues
Wood
- Primary fuel for 72% of the world population.
- Becoming scarce.
- Renewable with care & lower population size
Fossil Fuels
- Types
- Coal: may be high in sulfur.
- Oil: its cost drives economy.
- Natural gas: clean, cheap, but often from fracking.
- Problems: produce CO_2, Sox, particulates
Nuclear
- Conventional fission reactors
- U-234 split to release heat
- Heat produces steam
- Steam turns turbine
- Problems
- Expensive
- U.S. requires the latest available technology
- 3-12 times more cost overruns
- Cost without federal support is 13x oil
- Useful life is 40 years
- Shutdown is not defined
- No approved storage for waste: fuel rods are radioactive for > 100,000 years
- US has around 113 reactors in various parts of the country
- 67 were to be shut down by 2000
- Little new construction since 1977
- Fusion: hypothetical and expensive, but abundant & safe
Alternative Energies
- Hydropower
- Flowing water turns the turbine
- Few good places left to dam
- Tidal and river power are possibilities
- Solar
- Types
- Passive: south-facing windows
- Active: use fluid & pumps to move heat
- Thermal-electric: sun heats oil that boils water
- Photovoltaic: expensive
- Problems
- Sunshine availability
- Collector price and maintenance
- Wind
- Wind turns the turbine
- Clean, cheap
- Scarcity of sites
- Geothermal
- Let earth heat water
- Turn turbines
- But very few sites
- Biomass
- Grow crops for fuel (rice, cane, grasses)
- Burn or extract methanol
- Should balance its own CO_2 production
- Conservation
- Amounts to $10 billion per year
- Inexpensive
- But habits are hard to change once they are established
Future of Energy (at 2010 use, and proven reserves)
- Coal: 80% gone by 2100 AD
- Petroleum: by 2050 AD (or 2100 at the latest)
- Natural gas: by 2080 AD or perhaps 2110
- Nuclear: by 2200 AD
- Renewables (alternates)
- Infinite timespan
- But limited to about 35% of total needs
- Remember: as we use up one, we’ll rely more on others, and that will accelerate its depletion…
Population
General
- Populations naturally expand exponentially
Human
- Second billion in 1930
- Fourth billion in 1975
- Sixth billion in 1996
- Currently, 8.09 billion on the planet (April 2025)
- Doubling time
- Time required to double the population size
- Now is 42 years (1.7% growth rate)