Fungus

Kingdom Fungi:

Mycology: study of Fungi

Appeared about 400 mya

Not plants or animal

True multicellular with different types of cells

Strictly heterotrophic

Release digestive enzymes into environment that break down matter in their environment: extra cellular digestion

Some can digest dead organisms: decomposers

Some parasitic: feeding on living organisms and causing disease

Some predatory: can attack small worms in soil

Most saprophytic: decompose nonliving organic matter

Ex: yeast flour

In forests: leaves and animal waste

Along with bacteria, fungi are an important decomposer

Some decompose living tissue

Eg: athletes foot and ringworm

We use fungi to produce food and medicine

Biology of Fungi:

Body of fungus is mass of filaments called a mycelium

Each filament is a hypha

Hyphae allow organism to increase surface area in order to maximize contact with ground

Some have cross walls between cells called septa

Have pores that allow fungal cells to exchanges components of cytoplasm

Non septate fungi lack cross walls between cells: multinucleate

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Fungal cell structure:

Cell wall contains chitin instead of cellulose

Energy reserve is glycogen like in animal cells

Nonmotile

Move toward food source by growing towards it

Absorption heterotrophs: absorb nutrients from environment

Exhibit nuclear mitosis

Different from other eukaryotes

Chromosomes dragged to opposite poles by spindles inside nucleus

Process creates 2 cells

Fungal reproduction

Adapted to dry land

Product windblown spores

Spore: reproductive cell or multicellular structure that is resistant to environmental conditions and can develop into an adult without fusion with another cell

Haploid reproductive cell germinates and develops into new mycelium (asexual reproduction)

Sexual reproduction occurs by conjugation of 2 different mating types

Genetically different nuclei

Designated by + and -

After fusion of nuclei a zygote forms

Zygote undergoes meiosis

Produce haploid spores and cycle begins again

Fungi classified on basis of sexual structures

Phylum Zygomycota

Zygospore fungi

Saprotrophs and parasites

Eg: Rhizopus stolonifer: black bread mold

Phylum Ascomycota

Sac fungi

Sexual reproductive structure called ascocarp

Many sac fungi reproduce by producing chains of spores: conidia

Ex: disease causing sac fungi

Chestnut blight, dutch elm disease, Ergot

Candida causes thrush in humans

Some sac fungi useful to humans

Original source of penicillin

Used to produce blue cheese

Yeasts single celled fungi used in food industry

Morels and truffles

Phylum Basidiomycota

Club fungi

Ex: mushrooms, shelf fungi and puffballs

Structure includes stalk and cap with gills

Spores located in gills

Few edible species

Many toxic to humans

Rusts and smuts cause disease in plants and become a serious threat to food supply

Characteristic sexual reproductive structure: basidium

Contained within a basidiocarp

Edible part of a mushroom is basidiocarp

Located in gills of cap

Forms after fusion of + and - hyphae

Sexual reproduction is most common in this group

Asexual reproduction can occur by asexual spores

Lichens

Symbiotic relationship between fungi and cyanobacteria or green algae

Efficient at acquiring nutrients and moisture

Can colonize poor soil and rocky surfaces

Produce organic matter and create new soil

3 characteristic forms

Crustose (compact)

Foliose (shrub like)

Fruticose (leaf like

Body has 3 layers: fungi form top and bottom layers and protect middle layer of photosynthetic cells

Fungi portion offers protection and delivers water

Photosynthesis gives nutrients

Mycorrhizal fungi

Mutualistic relationship with plant roots

Help plants grow more successfully in poor soils

Fungi can live on outside surface of roots or can penetrate root tissues

Plant provides nutrients to fungus

Fungus bring water and minerals to plant

Hyphae provide lots of surface area for water absorption

Fungal disease of plants

Many enter through stomata of leaves or through a wound

Smuts and rusts: sac fungi that parasitize cereal crops

Fungal Disease of Humans (mycosis)

Athlete’s foot

ringworm

redness (inflammation due to enzymes released by fungus); extends outward in ring shape

Candidiasis (Candida albicans)

Yeast infection resulting from imbalance of normal flora

Histoplasmosis (Histoplasmosis capsulatum)

Can be transmitted by bird droppings

May be asymptomatic or mild flu like symptoms

In more serious cases: lesions may form in lungs which later become calcified

Control of Fungi:

Fungi more closely resemble animal cells than bacteria

Makes it harder to develop antibiotics that will kill fungi and not the host

Fungi synthesizes steroids differently

Fungicides are directed at steroid biosynthesis