7.2
Energy Transfer in Redox Reactions
Redox reactions are responsible for the transfer of energy throughout biochemical processes.
Electrons are carriers of energy.
Hydrogen atoms (which include electrons) can also transfer energy.
Role of NAD+ in Cellular Respiration
NAD+ serves as an electron carrier in metabolic processes.
It accepts two electrons along with a proton, transforming into NADH.
Role of Oxygen
Oxygen acts as a terminal electron acceptor in cellular respiration.
ATP as an Energy Source
ATP (Adenosine Triphosphate) is the most commonly used energy source in biological systems.
Breaking down ATP releases energy, which can fuel other reactions:
When coupled with an endergonic reaction (which requires energy), an exergonic reaction (which releases energy, such as ATP breakdown) can provide the necessary energy.
Substrate-Level Phosphorylation
Substrate-level phosphorylation involves the transfer of a phosphate group to ADP (Adenosine Diphosphate) to form ATP.
Phosphorylation: adding a phosphate group to ADP results in ATP formation.
Overview of Cellular Respiration
Cellular respiration consists of four key reactions, with the first two being:
Glycolysis:
Inputs glucose as a substrate.
Main Products:
2 Pyruvate molecules
2 NADH molecules
Location: Cytoplasm
The pyruvate produced from glycolysis moves into the mitochondria for further processing.
Pyruvate Oxidation
Pyruvate is oxidized to form Acetyl CoA.
Produces a by-product of CO2, which is considered a waste product.
The product that progresses to the next step is Acetyl CoA.
Mitochondrial Structure and Locations
Important areas within mitochondria for cellular respiration include:
Outer Membrane: Forms a barrier around the mitochondria.
Inner Membrane: Contains proteins involved in the electron transport chain.
Intermembrane Space: The space between the inner and outer membrane.
Matrix: Contains enzymes for the Krebs cycle and pyruvate oxidation.
Krebs Cycle (Citric Acid Cycle)
Acetyl CoA enters the Krebs cycle, which consists of nine reactions.
The process begins with Acetyl CoA combining with oxaloacetate to form citrate.
Rearranging citrate and decarboxylating two carbons release them as CO2.
Outputs for each Acetyl CoA:
2 CO2 (waste)
3 NADH (electron carriers)
1 FADH2 (another electron carrier)
1 ATP (via substrate-level phosphorylation)
Total yield of the Krebs cycle per glucose (accounting for two Acetyl CoA):
4 ATP
6 NADH
2 FADH2
Electron Transport Chain (ETC)
Definition: A series of protein complexes located in the inner mitochondrial membrane through which electrons are transported.
Function: Extracts energy from NADH and FADH2:
Electrons flow through protein complexes, pumping protons (H+) from the mitochondrial matrix into the intermembrane space.
Oxygen serves as the terminal electron acceptor, combining with protons to form water (H2O).
If oxygen is not present, the chain ceases to function.
Chemiosmosis
Involves protons flowing back into the mitochondrial matrix through ATP synthase:
ATP synthase works like a turbine, producing ATP as protons flow through.
This process is described as chemiosmosis.
ATP Yield in Cellular Respiration
Total potential ATP production from one glucose molecule through cellular respiration can vary:
Theoretical maximum yield is approximately 36 ATP.
Actual yield is often lower due to inefficiencies; typical yield is closer to 30 ATP.
The large majority of ATP is produced during the electron transport chain and chemiosmosis.
Summary of ATP Production Steps
Energy yield from glycolysis and Krebs cycle feeds into the electron transport chain.
Thus, understanding these processes alongside mitochondrial structure supports comprehension of cellular energy production during respiration.
Questions
Encourage questions on discussed topics, especially focusing on the roles and relationships in cellular respiration.