biol 3410 11/14 lec
Signal Integration
Introduction to signal integration in the nervous system.
Examining presynaptic neuron activity effects on postsynaptic neuron membrane potential.
Case Study of Presynaptic Neurons
3 presynaptic neurons; simultaneous firing leads to combined effects.
Example calculation based on EPSPs and IPSPs contributions to postsynaptic potential:
2 EPSPs: +10 mV and +12 mV, summing to +22 mV
1 IPSP: -5 mV, resulting in +17 mV overall.
Starting from -70 mV:
Final postsynaptic potential = -70 + 17 = -53 mV.
Action Potential Generation
If the resulting membrane potential is more positive than the threshold potential, an action potential is generated.
Clarification that -53 mV is above the threshold, thus an action potential will occur.
Types of Integration
Spatial Summation: Defined by simultaneous inputs from multiple neurons on a single postsynaptic neuron.
Temporal Summation: Involves rapid firing of a single presynaptic neuron, stimulating the postsynaptic neuron.
Often, signal integration in biological systems is a combination of both methods.
Spinal Cord Anatomy
Gray Matter:
Forms a butterfly shape in spinal cord; contains unmyelinated fibers and cell bodies.
White Matter:
Contains myelinated fibers, allowing for faster signal transmission.
Myelination Significance
Myelinated fibers found in spinal cord areas appearing white.
Gray matter consists of cell bodies and unmyelinated fibers, commonly linked to gray matter functions in reflex action.
Dorsal and Ventral Roots
Dorsal Roots: Carry sensory information; associated with sensory neurons.
Ventral Roots: Carry motor information; associated with motor neurons.
Spinal Nerves: Comprised of both dorsal and ventral roots merging.
Meninges
Protective coverings of spinal cord and brain include:
Dura mater (superficial)
Arachnoid mater
Pia mater (deep)
Reflex Arcs
Basic Components:
Receptor, sensory neuron (afferent), integrating center (spinal cord), motor neuron (efferent), and effector.
Responses can be either innate reflexes (like suckling) or learned reflexes (like driving).
Types of Reflexes
Monosynaptic Reflex Arc: Involves a single synapse (e.g., patellar reflex).
Polysynaptic Reflex Arc: Involves multiple synapses (e.g., withdrawal reflex when stepping on a nail).
Typically includes an interneuron that integrates the signal.
Example of Reflex Action
Patellar Reflex:
Triggered by tapping patellar tendon, stretching the associated muscle (quadriceps).
Action Potential travels through sensory neuron to spinal cord, activating motor neuron to cause muscle contraction.
Monosynaptic, Ipsilateral Reflex: Single pathway, same side of the body.
Neurological Mechanism in Reflex Action
The process involves the sensory neuron detecting stretch, synapsing with a motor neuron that causes muscle contraction while also suppressing the opposing muscle (hamstrings).
Neurotransmitters:
Peripheral muscle responses typically involve acetylcholine, while interneurons may utilize GABA for inhibition.
Summary
Signal integration through combined excitatory and inhibitory inputs is essential for neural function.
Reflex arcs facilitate rapid responses to stimuli through organized pathways, and the architecture of spinal components supports these processes efficiently.