Amniota - BIO205
Phylum Chordates (Chordata)
Definition: Animals with a notochord at some stage in their life cycle.
Sub-phylum Vertebrates (Craniata)
Vertebrates are chordates with vertebrae and a cranium.
Clade Gnathostomata: Vertebrates with jaws.
Clade Osteichthyes: Bony fishes and tetrapods.
Clade Tetrapods: Four-limbed vertebrates.
Clade Amniotes: Tetrapods with embryos having extraembryonic membranes.
Classification Overview
Protochordata:
Includes lancelets and tunicates.
Agnatha:
Includes hagfishes and lampreys.
Chondrichthyes:
Includes sharks, rays, and chimaeras.
Actinopterygii:
Ray-finned fishes.
Lobe-finned Fishes:
Ancestors to tetrapods.
Reptilia:
Includes turtles, lizards, snakes, crocodiles, and birds (Aves).
Mammalia:
Mammals characterized by hair and mammary glands.
Characteristics of Amniotes
Adaptations for a terrestrial lifestyle:
Amniotic Egg:
Contains four extra-embryonic membranes that support development.
Rib Ventilation of the Lungs:
More efficient method of breathing allowing greater activity levels.
High Pressure Cardiovascular System:
Facilitates increased metabolism via efficient oxygen delivery.
Thicker and More Waterproof Skin:
Protects against desiccation and environmental factors.
High Performance Kidneys:
Water-conserving kidneys to adapt to dry environments.
Expanded Brain and Sensory Organs:
Enhanced sensory processing and motor control.
Stronger Jaws:
Improved feeding mechanisms over ancestral forms.
Detailed Features of Amniotic Egg
Components of the Amniotic Egg:
Chorion:
Surrounds the embryo, acts as a respiratory organ.
Allantois:
Stores metabolic wastes, involved in gas exchange.
Amnion:
Protects the embryo by providing a fluid-filled cavity.
Yolk Sac:
Supplies nutrients to the developing embryo.
Mineralized Shell:
Provides mechanical support, limits water loss, and allows gas exchange.
Respiratory Adaptations
Lungs Development:
Increased surface area with the development of alveoli for gas exchange.
Rib ventilation improves lung capacity and efficiency.
Evolutionary examples: Salamander, Frog, Lizard, Mammal.
Cardiovascular Adaptations
High Pressure System:
Justifies double circuit circulation
Completely separated (in mammals, birds, and crocodiles).
Partially separated (in other reptiles).
Skin Adaptations
Thicker and Waterproof Skin:
Includes keratinized structures for protection (scales, hair, feathers).
Adaptations for moisture retention in terrestrial environments.
Sensory and Neurological Advancements
Expanded Brain and Sensory Organs:
Significant development in the cerebrum and cerebellum.
Enhanced processing of sensory information and muscle control for locomotion.
Conclusion
Amniotes demonstrate a suite of adaptations that allow them to thrive in diverse terrestrial environments, leading to evolutionary success in reptiles, birds, and mammals.