Type I-IV Hypersensitivity Reactions
Type I Hypersensitivity
- First exposure to pollen:
- Pollen binds to mast cells.
- No immediate symptoms.
- The state persists for a while.
- Re-exposure to pollen:
- Pollen interacts with immunoglobulin E (IgE) antibodies embedded in mast cells.
- Triggers degranulation of mast cells, releasing chemical mediators.
- This response essentially represents an acute inflammatory response.
- Chemical mediators released:
- Vasoactive amines (e.g., histamine).
- Lipid mediators (e.g., prostaglandins).
- Cytokines.
- Rapid effects of these mediators:
- Vasodilation.
- Increased vessel wall permeability (leading to swelling).
- Smooth muscle contraction in airways and lungs (in severe cases).
- Late phase reaction:
- Mast cells trigger later cytokine production.
- A delayed response that occurs some time after the initial reaction.
- Immediate reaction:
- Pollen binds to IgE receptors on mast cells.
- Triggers degranulation and release of histamine and other chemical mediators.
- Release of membrane phospholipids (arachidonic acid), leading to prostaglandins and leukotrienes.
- Effects of released substances:
- Prostaglandins and histamine: immediate vasodilation, increased vascular permeability, smooth muscle contraction.
- Leukotrienes and cytokines: recruit leukocytes, causing ongoing tissue damage via the inflammatory response.
- Histamine is rapidly inactivated by histaminase, limiting its acute effects.
- Severe reactions can be fatal.
- Late phase reaction: occurs several hours after exposure, less severe than the initial phase.
- Localized reactions:
- Urticaria (hives): welts on the body.
- Atopic dermatitis: a common form of dermatitis, often starting in childhood.
- Angioedema: swelling of the lips, tongue, etc.
- Systemic reactions (anaphylactic shock):
- Involve respiratory distress due to constricted bronchioles.
- System-wide vasodilation and increased vascular permeability can lead to cardiovascular failure and collapse.
- Symptoms include loss of consciousness, hives, tongue swelling, inability to swallow, rapid throat swelling.
- Urticaria (Hives):
- Characterized by a "wheel and flare" reaction.
- Wheel: central raised area with white oedema (swelling).
- Flare: surrounding redness due to vasodilation and inflammation.
- May be accompanied by sweating and a feeling of being hot and flushed.
- Angioedema:
- Extreme swelling due to increased vascular permeability in localized areas of the body.
- Urticaria triggers:
- Varied and can be unknown.
- Examples:
- Exposure to air.
- Heat or cold.
- Contact with a chemical allergen.
- Clinical settings:
- Treatments or products can trigger reactions.
- First aid training is essential.
- Monitor localized reactions for systemic symptoms.
- Do not let someone go home until symptoms improve. Escalate to ambulance if symptoms worsen.
- Anaphylactic shock:
- Adults with known allergies often carry an EpiPen for self-injection of adrenaline.
- Adrenaline counteracts histamine effects quickly, alleviating immediate symptoms.
- Buys time to get medical help.
Type II Hypersensitivity: Antibody-Dependent Cell-Mediated Cytotoxicity
- Involves the body responding to its own components, mediated by antibodies.
- Antigen is located on the cell surface or in connective tissue.
- Antibodies initiate an immune response against the body's own cells, leading to cell lysis, phagocytosis, or apoptosis.
- Also called antibody-dependent cell-mediated cytotoxic response.
- Mechanisms involved:
- Antibody binding activates the complement system.
- Complement proteins trigger a cascade, activating immune cells and intensifying the response.
- Complement cascade is initiated by a complement protein binding to the antigen-antibody complex on the cell.
- This leads to activation of further complement proteins.
- Complement fragments attract immune cells like neutrophils.
- Neutrophils bind to the antibody via Fc receptors on their surface.
- After binding, neutrophils release aggressive substances (e.g., radical oxygen species) that cause cellular damage, inflammation, and cell death.
- Activated complement proteins bind to the target cell surface, marking it for opsonization.
- Marked cells are recognized by phagocytes (e.g., macrophages).
- Macrophages use Fc receptors to bind to the antibody and engulf the target cell.
- The complement system leads to the formation of a membrane attack complex (MAC).
- MAC inserts into the plasma membrane, causing an influx of extracellular fluid.
- The cell swells and eventually lyses.
- Bound IgG antibodies are recognized by natural killer (NK) cells.
- NK cells possess Fc receptors that bind to the antibody, leading to NK cell activation.
- NK cells release perforin, which inserts into the plasma membrane, causing target cell lysis.
- NK cells also release granzymes, which initiate apoptosis.
- In some diseases, antibodies may be directed towards cell surface receptors, disrupting normal cell function.
- Antibody blocks the receptor, preventing signal transmission.
- Antibody binds to and activates the receptor.
- Type II hypersensitivity reactions usually cause cell death: complement-mediated or complement-independent.
- Cell death via lysis enhances inflammation.
- IgG and IgM antibodies are directed at cells of the body, causing damage through:
- Complement actions (opsonization, phagocytosis).
- Recruiting phagocytes.
- Complement-mediated inflammation (membrane attack complex).
- Changes to receptors.
- Cell dysfunction in thyroid cells: antibodies against thyroid cell receptors turn on thyroid hormone production.
- Antibodies binding to receptors can also block receptor action.
- Pemphigus vulgaris:
- Activation of a protease that cleaves desmosome connections between keratinocytes.
- Leads to a blistering condition.
- Keratinocytes split apart, causing exudate and inflammation.
Type III Hypersensitivity: Immune Complex Reactions
- Certain ratios of antigens and antibodies form complexes that are too small to be phagocytosed.
- These complexes get trapped in tissues, triggering complement-mediated inflammation.
- Recruits inflammatory cells, causing tissue damage.
- Often occurs in blood vessels or connective tissue.
- Diseases have a range of symptoms.
- Systemic lupus erythematosus:
- Can present as a mask-like rash on the face.
- Also has constitutional symptoms (e.g., joint issues).
- Immune complexes consist of antibodies bound to antigens.
- Antigens and immune complexes float freely in the blood (soluble immune complexes).
- Antigens can be external (foreign) or internal (self).
- IgM or IgG antibodies bind to antigens, forming complexes of various sizes.
- Smaller immune complexes are more likely to remain in blood circulation.
- Circulating immune complexes deposit on the blood vessel wall, especially in small vessels.
- Immune complexes rapidly initiate the complement system.
- Neutrophils are attracted by activated complement proteins and bind to the antibody, releasing destructive substances.
- Damage to endothelial cells increases vessel permeability, resulting in leakage into surrounding tissue, causing inflammation and damage.
- Discriminating features from Type II hypersensitivity:
- Type III initiating components are soluble before depositing on the vessel wall.
- Tissues with small blood vessels are most affected.
- Complement system activation is more rapid and stronger in Type III.
- Immune complex lodges within vessels, recruiting neutrophils, triggering inflammation.
- Acute inflammatory response can become a chronic inflammatory response.
- Lupus:
- Kidneys and joints are affected.
- Fine vasculature of vessels in the face is affected.
- Leads to a butterfly-type rash.
Type IV Hypersensitivity: Delayed-Type Hypersensitivity
- Harmful effects produced by cell-mediated immunity.
- Mediated by Th1 cells (TD cells).
- Responsible for contact dermatitis, tissue damage in infectious diseases, and tissue graft rejection.
- Contact hypersensitivity (e.g., poison oak dermatitis):
- Initiated by contact with the antigen (allergen).
- A hapten binds to a carrier protein in the host.
- The complex is ingested by a macrophage and presented on a class II MHC.
- Th1 cells recognize the antigen, become activated, and increase in numbers.
- Second exposure:
- Th1 cells react with the antigen. Release cytokines. This results in attraction of more macrophages followed by inflammation and skin lesions.
- Characteristic skin lesions appear after 24 hours, peaking at 48-72 hours.
- Timeline: delayed hypersensitivity due to the timeline for reaction. This means it could appear 12-72 hours after exposure.
- Antigen presenting cells, like Langerhans cells in the skin, uptake allergens.
- They process and present it with its major histocompatibility type two molecule, and they migrate to lymph nodes, where they can then interact with helper T cells and CD8 positive T cells to trigger those reactions within tissues.
- Inflammatory and cell lysis responses.
- CD4 T cells produce pro-inflammatory cytokines, leading to tissue injury.
- CD8 positive or cytotoxic T cells respond, producing porphyrins or branulycin or granzymes that damage local tissue.
- Direct action of the CD8 positive T cells causing that damage directly without necessarily the inflammatory component of that.
- Diseases include contact dermatitis, type one diabetes, rheumatoid arthritis, multiple sclerosis, inflammatory bowel disease, and coeliac disease.
- Contact dermatitis:
- Rash develops on areas of contact.
- Appears as a dry, irritated dermatitis.
- Common allergen is the adhesive on dressings.
- Localized around the lips: suspects could be food, drink, makeup, or skincare products.
- Around the eyes: could be from products that have come into contact.
- Nickel allergies: rashes on areas of contact with cheap metal jewellery.
Photo Reactions
- Aberrant reaction where the allergen (photoallergic) or toxin (phototoxic) alone cannot cause the reaction.
- Certain wavelengths of light (usually UVA and visible light) trigger the reaction.
- Photo reactions come in two types: photo irritancy and photo allergic.
Phototoxic Reaction (Photoirritancy)
- Chemical substance, when exposed to light, causes direct damage or direct inflammatory damage to tissues.
- No allergic response.
- Appears like a bad sunburn, often with blistering.
- Post-inflammatory hyperpigmentation is common.
- Known photosensitizing chemicals will cause a reaction in pretty much anyone exposed to those.
- Due to a direct tissue response not specific to individual.
- Phototoxic molecules in a low excitability state change to a high excitability state when exposed to light.
- Highly excitable molecules convert neighboring molecules to free radicals, causing tissue damage and triggering inflammation.
- Over time, can develop into a photoallergy as well.
- Common photo toxic compounds include:
- Medications: antibiotics (tetracycline, sulfonides, and quinolones), antifungals (Resifulvan), other pharmaceuticals (Sorrelin's five fluorouracil, naproxen).
- Topical agents: coal tar preparations, essential oils (citrus oils).
- Plants: parsley.
- Naturally occurring molecules: porphyrins.
- Dermal clinicians:
- Use light-based devices and can trigger these reactions.
- Thorough client consultation is needed to check for exposure to known phototoxic agents (medications).
- MEMS database (drug database) explains the properties of different medications, including their phototoxic potential.
- Photosensitizing medications are often a red flag contraindication for treatments.
Photoallergic Response
- Requires light and a chemical compound in someone who is allergic.
- A type IV reaction
- It looks a lot more like the allergic contact dermatitis.
Comparing Photo Reactions: - Phototoxic vs. Photoallergic:
- Often difficult to differentiate.
- Phototoxic: quicker, requires larger exposure, develops only on sun-exposed skin, looks like an exaggerated sunburn with blistering.
- Photoallergic: has a rough drier looking erythematous rash like the type one reactions the type three reactions rather. Longer time. 24-72 hours to develop. Type IV immune response reaction.
- Hyperpigmentation
* Generally speaking it's what we call post inflammatory hyperpigmentation. So it is in most instances fairly superficial. And in most cases that would clear up in six months or so.
* It's a shorter term issue.