Techniques in Cell and Molecular Biology
Chapter 18: Techniques in Cell and Molecular Biology
Microscopy
Looking at Cells
Light Microscope Components:
Ocular lens
Light beam
Objective lens
Specimen
Condenser lens
Light source
Light Microscope Description:
Utilizes glass lenses and visible light to form an image.
Resolution of approximately 0.2 ext{ } ext{μm}; 1000 times greater than the human eye.
Visualizes cell sizes, shapes, and some internal cell structures.
Internal structures are difficult to see under visible light; involves chemical treatment and staining with various dyes to enhance contrast.
Reference: LIFE: THE SCIENCE OF BIOLOGY 11e, Figure 5.3 (Part 1)
Specialized Light Microscopes
Techniques Developed to Observe Living Cells:
Phase-Contrast Microscopy:
Increases contrast by emphasizing differences in refractive index, enhancing light and dark regions.
Differential Interference Contrast (DIC) Microscopy:
Uses two beams of polarized light to create images that appear as if the cell casts a shadow.
Fluorescence Microscopy:
A method to visualize structures using fluorochromes that emit light when excited by UV rays.
Confocal Microscopy:
Produces sharp images of thin sections, utilizes laser and pinhole to eliminate out-of-focus light.
Bright-Field Microscopy
Characteristics:
Light passes through specimen; if no natural pigments exist, it presents low contrast.
Image detail may be enhanced by staining.
Staining and Contrast Enhancement
Stains Variety:
Stains chemically differ and bind to different cell materials. They significantly improve visualization in microscopy.
Fluorescence Microscopy Techniques
Immunofluorescence:
Uses fluorochrome-conjugated antibodies to study specific cellular components.
Green Fluorescent Protein (GFP):
Isolated from jellyfish Aequorea victoria by Osamu Shimomura in the 1960s.
GFP can be fused with target genes in transgenic model organisms for live-cell imaging and gene expression tracking.
Applications include studying protein localization in living cells, following dynamic neuronal interactions.
GFP Variants:
Developed by Roger Tsien; used to create brighter and color-shifted fluorescent proteins.
Impact of GFP:
Revolutionized molecular biology by enabling live-cell imaging.
Awarded the Nobel Prize in Chemistry in 2008 to Shimomura, Chalfie, and Tsien.
Examples of GFP Applications
GloFish:
Transgenic zebrafish containing red fluorescent protein useful in environmental testing.
GFP in Research:
Used to observe gene expression and protein interactions under various conditions.
Fluorescent In Situ Hybridization (FISH)
Technique Description:
Allows detection of specific nucleic acid sequences within fixed cells.
Application:
Gene mapping and visualization of chromosomal regions.
Immunostaining Techniques
Methods to Detect Proteins:
Uses primary antibodies binding to specific antigens, secondary antibodies for visualization.
RNA-FISH Techniques
RNA Detection Methodology:
Involves dissection, fixation, prehybridization, and imaging subsequent to hybridization with fluorescent probes.
Advanced Fluorescence Techniques
Fluorescence Resonance Energy Transfer (FRET):
Measures distances between molecules using different fluorescent labels to detect protein-protein interactions.
Confocal vs Regular Fluorescence Microscopy Comparison
Regular Fluorescence Microscopy:
Broad sample illumination, captures both in-focus and out-of-focus light; quicker but can cause blurred images.
Confocal Microscopy:
Laser focus on thin optical sections, produces clearer images, and can construct 3D representations.
Electron Microscopy (EM)
Types of Electron Microscopy:
Transmission Electron Microscopy (TEM):
Uses electrons transmitted through the specimen; higher magnification up to 1,000,000×.
Scanning Electron Microscopy (SEM):
Generates detailed 3D images from electrons reflected off the specimen's surface.
Freeze-Fracture Replication:
Technique involves freezing specimens, fracturing them, and visualizing their internal structures in TEM.
Atomic Force Microscopy (AFM)
Description:
High-resolution technique for obtaining topographical images of individual molecules.
Applications:
Real-time imaging and manipulation of macromolecules, receptor/ligand interactions measurement.
Cell Culture Techniques
Definition:
The practice of growing cells under controlled conditions for biological studies.
Types of Culture:
Primary culture (cells direct from organisms), Secondary culture (derived from a primary culture), Cell lines (immortal cells).
Studying Organelles and Cell Fractionation
Methods for Studying Organelles:
First using light microscopy, then electron microscopy, staining techniques targeted to identify organelle contents.
Cell Fractionation:
Separates organelles based on size/density via centrifugation.
Protein Purification Techniques
Process Description:
Involves removing contaminants to enhance specific protein activities.
Methods:
Liquid column chromatography; separates components via affinity to a material; High-performance liquid chromatography (HPLC) for resolution.