Bacteria 1 The Streptococci MID 2029
Learning Objectives
Describe the medically pertinent anatomy of Streptococci:
- Membrane
- Cell wall
- Gram-stainList the streptococci by genus/species and by Lancefield group (if appropriate)
List any characteristic morphologies of the various streps:
- Chains
- Pairs
- Lancet-shaped
- CapsuleBe familiar with immunizations for any species of Streptococcus:
- Indications
- TargetsOrganize the clinical diseases associated with each Strep species
Associate Streptococcus pyogenes (Lancefield group A) with its immunologic implications in heart, kidney, and systemic diseases.
Order the Strep species by antibiotic resistance, from least to most resistant.
Strep Vocabulary List
Gram-positive cocci
Blood agar
Hemolysis
Catalase negative
Peptidoglycan
Antibiotics:
- Penicillin
- Cephalosporin
- VancomycinStreptococcus pyogenes:
- Lancefield group A
- Also known as flesh-eating strep
- Associated diseases include cellulitis, pharyngitis, scarlet fever, erysipelas, pyoderma, impetigo, necrotizing fasciitis
- Erythrogenic toxinASO (Antistreptolysin O) titers
M protein
Rheumatic fever
Post-streptococcal glomerulonephritis
Toxic shock syndrome and superantigens
Streptococcus agalactiae:
- Lancefield group B
- Causes pneumonia, sepsis, meningitis in neonatesStreptococcus pneumoniae:
- Also known as pneumococcus, Strep pneumo
- Immunizations available (polysaccharide capsule)
- Morphology: lancet-shapedEnterococcus:
- Causes urinary tract infections, sepsis, subacute bacterial endocarditis (SBE)
- VRE (vancomycin-resistant Enterococcus)Viridans Strep:
- Includes Streptococcus mutans and mitis
- Also includes Streptococcus sanguinis.
The Why Question
Consider the prevalence of Strep throat:
- Strep infections can lead to:
- Rheumatic fever affecting the heart
- Post-streptococcal glomerulonephritis which can cause renal failureStreptococcus pneumoniae is a significant cause of pneumonia and meningitis.
Streptococcal toxic shock syndrome led to the death of Jim Henson, the creator of the Muppets.
Curiosity
Spend a few moments contemplating questions you might have about bacterial infections, Streptococcus, and immunology.
Write these questions down and seek the answers throughout this presentation.
Your curiosity is essential in becoming a life-long learner.
Resources for Further Learning
Murray Microbiology, especially the first couple of pages of each chapter
First Aid (2025 or 2026 edition)
Robbins chapter on Infectious Diseases (chapter 8)
Previous year’s notes from Dr. Gomez.
Streptococci Overview
General Characteristics:
- Gram-positive cocci typically arranged in chains (with the exception of S. pneumoniae which is lance-shaped and diplococci)
- Catalase-negative
- Non-motile
- Either facultative or obligate anaerobesLancefield Groupings:
- Based on serology of cell wall C carbohydrate (total of 18 groups)Virulence Factors:
- M proteins in S. pyogenes resist phagocytosis
- Lipoteichoic acids facilitate attachment to epithelial cells
- S. pneumoniae, S. pyogenes, and S. agalactiae possess a capsule:
- S. pyogenes has a hyaluronic acid capsule
- Major toxins:
- Streptolysins
- NADase
- Hyaluronidase
- Streptokinase
- DNAses
- Erythrogenic exotoxin (causes rash in scarlet fever)
- Toxic shock syndrome toxin
- Pyrogenic exotoxin B (in erysipelas)
Group A Streptococcus (S. pyogenes)
Hemolysis and Classification:
- β hemolysis: primary cause of bacterial pharyngitis, tonsillitis, sinusitis, otitis, arthritis, osteomyelitis, impetigo, cellulitis.
- Can later cause rheumatic fever, rheumatic valvular disease, and post-streptococcal glomerulonephritis.
- Can also cause scarlet fever (due to toxin) and pneumonia, bacteremia, necrotizing fasciitis, and toxic shock syndrome.
Group B Streptococcus (S. agalactiae)
Characteristics:
- β hemolytic
- Leading cause of neonatal septicemia, meningitis, and pneumonia
Group D Streptococcus
Includes S. bovis and S. equinus
ɑ or γ hemolytic
Associated with urinary tract infections and endocarditis
Streptococcus pneumoniae
Synonyms and Morphology:
- Known as pneumococcus, Strep pneumo, or friend of the elderly
- Gram-positive cocci in pairs (lancet-shaped diplococci)
- ɑ hemolytic
- Encapsulated with polysaccharide capsule, 94 types available for immunization.Diseases Caused:
- Primary causes respiratory infections including:
- Lobar pneumonia: Symptoms include cough, rusty sputum, fever
- Meningitis: Most common in adults.
- Bacteremia in individuals without a spleen or with sickle cell disease.
Epidemiology of S. pneumoniae
Carried in the nasopharynx in 5-50% of individuals
Common in children: severe otitis media, pneumonia, and meningitis.
Common in adults: pneumonia and meningitis.
Drug resistance: up to 20% of S. pneumoniae are resistant to beta-lactams and macrolides.
Immunization:
- Available for the capsule polysaccharide (PPS) or conjugate forms (PCV 15 or 20, PPSV 23).
More on Alpha Hemolytic Streptococcus
Viridans Strep:
- Associated with dental health (S. mutans and S. mitis): Causes dental caries.
- S. sanguinis: Associated with subacute bacterial endocarditis (SBE).
Beta Hemolytic Streptococcus
Characterized by type-specific antigens.
Lancefield Groups:
- Group A = Strep pyogenes, commonly causes sore throat
- Group B = Strep agalactiae
- Other groups include C and D which are less common.
Streptococcus pyogenes (Group A strep)
Characteristics and Diseases:
- Gram-positive cocci in chains
- Causes strep throat (pharyngitis), which can be complicated by erythrogenic toxin leading to Scarlet Fever.
- Skin infections include:
- Erysipelas (superficial infection)
- Pyoderma and impetigo (common in children, presents with a honey-like crust)
- Cellulitis and necrotizing fasciitis (deeper infections).
Identification and Laboratory Testing for S. pyogenes
Rapid strep test: immunologic typing of Group A.
Culture: typically produces beta hemolytic colonies—subsequent typing (A or B); sensitive to Bacitracin.
ASO (Anti-Streptolysin O) titers may also be used for diagnosis.
Treatment:
- Not highly resistant, so penicillin works well; alternatives include erythromycins/macrolides and vancomycin.
Virulence and Immunologic Implications of S. pyogenes
M proteins in the cell wall assist in avoiding phagocytes (PMNs).
Cross-reactivity with heart tissue:
- Leads to rheumatic fever manifesting in the heart, valves, and joints.Post-streptococcal glomerulonephritis results from immune complex deposition in the kidney.
Toxic shock syndrome toxin can be overwhelming, similar to staphylococcal toxic shock syndrome toxins known as superantigens.
Summary of Key Concepts
Characteristics of Gram-Positive Organisms:
- Gram-positive organisms have a thick peptidoglycan layer in their cell wall which retains violet stain, making them appear blue-black under a microscope.
- They typically have a single cell membrane.
Identification Guidelines
Strep species grow best on blood agar, a nutrient-rich medium that also serves as an indicator for hemolysis.
Distinguished from Staphylococci: Staphylococci produce catalase (catalase positive), whereas Streptococci are catalase negative.
Hemolysis Types:
- Alpha: Partial hemolysis – greenish
- Beta: Complete hemolysis – halo appearance
- Gamma: No hemolysis
Enterococci
Characteristics:
- Gram-positive diplococci or short chains (previously classified with group D streptococci)
- Catalase negative, γ hemolytic (non-hemolytic)
- Non-motile, facultative anaerobesDiseases:
- Can cause infections like urinary tract infections, bacteremia, bacterial endocarditis, diverticulitis, and meningitis.
- E. faecalis and E. faecium are part of the normal intestinal flora.
- These species show intrinsic resistance to beta-lactam antibiotics (penicillins, cephalosporins, carbapenems) and many aminoglycosides, leading to VRE (vancomycin-resistant Enterococcus).
Streptococcus bovis
An oddball gamma Streptococcus, lives in the colon but is associated with health complications, including endocarditis and sepsis frequently correlated with colorectal cancer (associated with subspecies gallolyticus).
Evaluations and Practice Questions
Utilize various presented scenarios to assess knowledge of Streptococcus, its pathogenesis, and treatment options.
- Discuss clinical manifestations presented in cases of glomerulonephritis, pneumonia, and maternal-fetal transmission and the relevance to antibiotics and immunization strategies.