Metabolism Overview
Metabolism Overview
What is Metabolism?
Definition of Metabolism: A set of chemical reactions in living organisms that sustain life.
Functions:
Facilitates growth.
Maintains homeostasis.
Enables reproduction.
Allows adaptation to the environment.
Types of reactions:
Anabolic Reactions: Build complex molecules from simpler ones.
Catabolic Reactions: Break complex molecules into simpler ones, releasing energy.
Major Metabolic Pathways
Substrates:
Fats: Broken down into fatty acids and glycerol.
Polysaccharides: Broken down into glucose and other sugars.
Proteins: Broken down into amino acids.
Stages of Metabolism:
Stage I: Fatty acids and other sugars are converted.
Stage II: Conversion to Acetyl CoA and entry into the Citric Acid Cycle.
Stage III: Electrons are transferred to O2 via oxidative phosphorylation, producing ATP and H2O.
The Truth About Metabolism
Commonalities:
All organisms share common metabolic pathways.
Limited number of molecules involved across pathways.
Shared classes of chemical reactions.
Simple mechanisms of reactions.
Similar process control points.
Conservation: What enters the system must eventually be expelled.
Catabolism
Characteristics:
Degradation reactions that produce energy.
Reduce molecular complexity.
Example Reaction: Convert fuels (carbohydrates, fats) into CO2, H2O, and energy.
Gibbs Free Energy ( \u0394G'):
Typically negative in catabolic reactions.
Examples include:
Carbohydrate degradation.
Amino acid degradation.
Fatty acid degradation.
Anabolism
Characteristics:
Synthesis reactions that require energy.
Create complexity and energy-rich substrates.
Reaction Example: Useful energy + Simple precursors = Complex molecules.
Gibbs Free Energy ( \u0394G'):
Generally positive in anabolic reactions.
Examples include:
Amino acid synthesis.
Fatty acid synthesis.
ATP and Energy Transfer
Key Components:
Simple molecules: Glucose, amino acids, and glycerol.
Energy storage and transfer through ATP.
Processes:
Catabolic reactions transfer energy from complex molecules to ATP.
ATP can be converted to ADP and inorganic phosphate (P).
Energy is also released as heat.
Oxidation and Reduction in Metabolism
Biological Oxidation: Involves loss of hydrogen atoms/electrons (dehydrogenation).
Reduction: Involves gain of hydrogen atoms/electrons, increasing energy potential.
Coenzymes that facilitate these processes:
NAD (Nicotinamide adenine dinucleotide)
NADP (Nicotinamide adenine dinucleotide phosphate)
FAD (Flavin adenine dinucleotide)
Carbohydrates
Definition: "Hydrates of carbon."
Types:
Monosaccharides, disaccharides, trisaccharides, oligosaccharides, and polysaccharides.
Functions:
Energy storage.
Provide fuel and metabolic intermediates.
Structural components of biomolecules.
Carbohydrate Structures
Monosaccharides: Characterized by carbon count:
Tri – 3 carbons.
Tetr – 4 carbons.
Pent – 5 carbons.
Hex – 6 carbons.
Hept – 7 carbons.
Structure: Mostly contain an OH group, and the last carbon has either an aldehyde or ketone group.
Fischer Projection
Characteristics:
Carbon chain represented vertically; horizontal lines indicate bonds projecting towards the viewer.
C1 is always at the top; cannot rotate within the plane of the page.
Creates enantiomers (non-superimposable mirror images).
Stereochemistry
Highest numbered stereocenter indicates the configuration (D or L designation):
If it resembles D-glyceraldehyde, it’s denoted as D.
Cyclic Sugars
Cyclic Structures:
Pyran and furan forms of sugars exist; examples include D-glucopyranose and D-fructofuranose.
Glycosidic Linkages
Formation of glycosidic bonds involves 'locking' the anomeric configuration.
Non-reducing and reducing sugars are characterized based on linkage types (example: lactose, sucrose).
Polysaccharides
Examples include:
Cellulose: Composed of beta-1,4 linkages.
Starch and Glycogen: Composed of alpha-1,4 linkages with branching.
Glycogen structure includes multiple linkage types (alpha-1,6 for branching).
Carbohydrate Metabolism Summary
Key processes:
Glycogenesis: Formation of glycogen from glucose.
Glycogenolysis: Breakdown of glycogen.
Pentose phosphate pathway: Converts glucose for nucleotide synthesis.
Gluconeogenesis: Formation of glucose from non-carbohydrate sources.
Glycolysis: Breakdown of glucose to pyruvate.
Citric Acid Cycle and Electron Transport Chain: Final ATP production.
Conclusions
Metabolism consists of energy-yielding and energy-requiring reactions.
Catabolic processes degrade macromolecules while yielding ATP.
Anabolic processes utilize energy to construct complex molecules from simpler ones.