Proteins

Chemistry of Proteins

Learning Objectives

  • Categorize amino acids based on functional group and polarity

    • Amino acids have functional groups like amino and carboxyl groups.

  • Explain the amphoteric property of amino acids

    • Due to the presence of amino and carboxyl groups.

  • Illustrate the formation of peptides from different amino acids

  • Determine amino acid sequences and peptide names

  • Importance of small peptides

  • Describe levels of protein structure and their interrelationship

  • Relate protein structure to physiological function

  • Differentiate hydrolysis and denaturation of proteins

Overview

    1. Protein: Crucial biomolecules in the body, derived from the Greek word proteios.

    2. Proteins: account for 15% of cell mass and half of its dry weight.

    3. Proteins: are unbranched polymers of amino acids linked by peptide bonds.

    4. Structure: Collagen and keratin provide structural support as fibrous proteins.

    5. Catalysis: Enzymes catalyze reactions in organisms for various functions.

    6. Movement: Proteins like myosin and actin enable muscle mobility.

    7. Transport: Hemoglobin transports molecules in blood and across membranes.

    8. Amino Acids: Building blocks of proteins containing carboxyl and amino groups.

    9. Essential Amino Acids: Body cannot synthesize them adequately; must be obtained from the diet.

    10. Standard Amino Acids: Humans have 20 standard amino acids with distinct side chains called.

    11. Stereoisomers: Standard amino acids have stereogenic centers, existing as D or L enantiomers.

    12. Fischer Projection: Representation method for D and L stereoisomers of amino acids.

    13. Side Chains: Distinguish amino acids and group them by the polarity of their side chains.

    14. essential amino acids: 10 amino acids crucial for a child's normal growth and development.

  • Arginine is essential for infants for normal growth but becomes nonessential as they grow.

  • Premature infants may lack sufficient quantities of some nonessential amino acids, making them conditionally essential until maturity.

  • Conditionally essential amino acids must be obtained through the diet, found in human milk and infant formula.

  • Complete dietary proteins contain all essential amino acids in adequate amounts.

  • Proteins from animal sources are usually complete, while incomplete proteins lack one or more essential amino acids.

  • Gelatin is an incomplete protein with tryptophan as its limiting amino acid.

  • Proteins from plant sources are generally incomplete, with lysine, methionine, and tryptophan as common limiting amino acids.

  • Soy protein is a complete plant protein, while a mix of plant proteins like rice and beans can provide complete dietary protein.

  • Eating rice and beans together forms complementary dietary protein.

Amphoteric Properties of SAAs

  • Amino acids have both acidic (-COOH) and basic (-NH2) groups in one molecule.

  • At neutral pH, -COOH loses a proton becoming carboxylate, while -NH2 accepts a proton becoming a quarternary ammonium ion.

  • In aqueous solutions, -COOH is deprotonated, and -NH2 is protonated in SAAs.

pH Effects on Amino Acid Structure

  • Amino acid structure varies at different pH levels.

  • In acidic solutions, carboxyl group ionizes first, while in basic solutions, the ammonium group ionizes first.

  • Different structures are observed at varying pH levels for acidic and basic amino acids.

Specific Ionization Patterns

  • Histidine, lysine, and arginine have distinct ionization patterns at different pH levels.

  • Histidine's imidazolium group ionizes first, while lysine and arginine have the quat. Ammonium group of the a-carbon ionizing first.

  • Acidic to basic solution pH solutions

    • Acidic amino acids are amphoteric, can be both acidic and basic

    • Carboxyl group can be acidic or deprotonated near neutral pH

    • Protonated NH2 becomes positively charged NH3

  • Conjugate acid pattern and conjugate base formation explained

  • Peptides

  • Peptide formation by condensing carboxyl group and amino group to form a peptide bond

  • Nomenclature of peptides

    • C-terminal amino acid residue keeps its full name

    • Other residues named with -yl suffix

    • Naming sequence starts at the N-terminal acid residue

  • Isomeric peptides contain similar amino acid residues but in different orders

  • Oxytocin and Vasopressin are small peptides with physiological importance

    • Functions and roles of oxytocin and vasopressin explained

  • Glutathione as an antioxidant peptide

  • Primary structure of proteins defined by the number, kind, and sequence of amino acids

    • Importance of primary structure in determining the protein's three-dimensional structure and function

  • Importance of knowing the primary structure of a protein

    • Primary structure provides the sequence of amino acids in a protein.

    • Helps in understanding the function and properties of the protein.

    • Example of amino acid sequences in different animals provided.

  • Secondary Structure of Proteins

    • Refers to the regular localized arrangement of the polypeptide backbone.

    • Stabilized by hydrogen bonds between C=O and N-H groups.

    • Examples include alpha-helix and beta-pleated sheet structures.

  • Factors disrupting the alpha-helix structure

    • Presence of proline, causing a break in the helix.

    • Adjacent similarly charged groups leading to electrostatic repulsion.

    • Adjacent bulky groups causing steric repulsion.

  • Tertiary Structure

    • Overall three-dimensional structure of proteins.

    • Results from interactions of side chains of amino acid residues and prosthetic groups.

  • Quaternary Structure

    • Non-covalent association of protein subunits into a supramolecule.

    • Not all proteins have quaternary structures.

  • Different quaternary structures based on the number of subunits

    • Examples: Dimer (Insulin), Trimer (Thrombin), Tetramer (Hemoglobin).

  • Protein Hydrolysis

    • Causes disruption of peptide bonds, liberating free amino acids.

    • Can be carried out using strong acids, bases, or enzymes (proteases).

  • Protein Denaturation

    • Leads to the unfolding of a protein's three-dimensional structure.

    • Results in the loss of biological activity.

    • Denaturation can be reversible or irreversible.

  • Factors causing denaturation

    • Denaturing agents like heat, urea, detergents, acids, bases, salts, reducing agents, heavy metals, and alcohol.