The study of how evolutionary processes shape inherited behaviors and the ways that animals respond to specific stimuli
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Behavior
An animal’s response to a stimulus (internal or external)
Example: nature vs. nurture (genetic and environmental factors) and allow for survival and reproduction (subject to natural selection)
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Proximate cause
How a behavior occurs or how it is modified
1. What was the stimulus to cause the behavior 2. How does the “nurture” component affect behavior? (How do the experiences during growth and development influence the response)
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Ultimate cause
Why a behavior occurs (in context of natural selection)
1. How does the behavior help the animal survive and reproduce? 2. How does the “nature” component affect behavior? (What is the evolutionary basis of the behavior)
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Innate behaviors
Developmentally fixed
* hereditary, born behaviors, do not need to learn them; instinctive * Experience during growth have no obvios effect
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Learned behaviors
Depend on environmental influence
* experiences do affect these behaviors * High variation in a population
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Fixed action patterns (FAPs)
Sequence of unlearned acts directly linked to a stimulus
* actions are unchangeable * Carried out to completion * Triggered by a sign stimulus (external cue)
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Migration
A regular, long distance change in location
* triggered by environmental cues (sun’s position, earth’s magnetic field and celestial cues)
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Signal
A stimulus generated and transmitted from one animal to another; animal communication
* examples: visual, auditory, tactile, electrical, chemical
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Pheromones
Chemicals emitted by members of a species that can affect other members of the same species
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Stimulus response chains
When a response to a stimulus serves as the next stimulus for a behavior
* seen in animal courtships
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Body movement
Example: waggle dance in bees
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Directed movements
Movements towards or away from a stimulus
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Kinesis
A change in the rate of movement or the frequency of turning movements in response to a stimulus; non directional
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Taxis
Directional movement towards (positive) or away from (negative) a stimulus
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Phototaxis
Movement in response to light
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Chemotaxis
Movement in response to chemical signal
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Geotaxis
Movement in response to gravity
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Learning
The modification of behavior based on specific experiences
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Imprinting
A long lasting behavior response to an individual
* happens during a sensitive period of development (usually very early in life) * Imprinting occurs on the first individual they encounter
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Spatial learning
Establishing memories based upon the spatial structure of the animal’s surroundings
* some animals form a cognitive map or use landmarks as environmental cues (example: birds finding their hidden nests)
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Associative learning
The ability to associate one environmental feature with another
* example: associating monarch butterflies with foul taste
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Social learning
Learning through observations and imitations of the observed behaviors
* example: chimps breaking open oil palm nuts
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Foraging
Food obtaining behavior
* searching for, recognizing and capturing food items * Animals better at foraging will be more successful in finding food
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Mating behaviors
Animals can be monogamous or polygamous (polygyny or polyandry)
* sexual dimorphism can result from sexual selection
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Altruism
Selfless behavior
* reduced the individual’s fitness, but increases the fitness of the rest of the population * Example: naked mole rat colonies have only one reproducing female (queen), who will only mate with a few males (kings). The other reproductive members will sacrifice themselves to protect their queen and kings
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Phototropism
A directional response that allows plats to grow towards (an in some cases away from) a source of light
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Photoperiodism
Allows plants to develop in response to day length; plants flower only at certain times of the year
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Response in plants
* Defend themselves against herbivory * physical defenses: thorns, trichomes (small plant like hairs) * Chemical defenses: production of toxic or distasteful compounds
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Soil composition
* soil composition can affect plants * The pH of soil can affect flower coloring in some plants * Nutrients are more accessible at certain pH * Example: hydrangea blooms turn different colors based on soil pH
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Ecosystem
the sum of all the organisms living in a given area and the abiotic factors they interact with
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Biotic factors
Living, or once living, components of an environments
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Abiotic factors
Non living (physical and chemical properties of the environment)
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1st law of thermodynamics
energy can neither be created nor destroyed, only transferred
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2nd law of thermodynamics
Exchanges of energy increase the entropy of the universe
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Net gain in energy
Net gain results in energy storage or growth of an organism
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Net loss of energy
Results in loss of mass and eventual death of an organism
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Metabolic rate
The total amount of energy an animals uses in a unit of time
* can be measured in calories, heat loss or by the amount of oxygen consumed (or CO2 produced) * An animal’s metabolic rate is related to its body mass
Use thermal energy from metabolism to maintain body temperature
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Ectotherms
Use external sources (ie. sun/shade or other organisms) to regulate their body temperature
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Tropic levels
Species can be grouped into tropic levels based upon their main source of nutrition and energy
* energy cannot be recycled * The sun constantly supplied energy to ecosystems
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primary producers
Use light energy to synthesize organic compound
* plants, algae, photosynthetic plankton
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Chemosynthetic
they produce food using the energy created by chemical reactions
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Heterotrophs
rely on autotrophs because they cannot make their out food
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Primary consumers
Herbivore
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Secondary consumers
Carnivores that eat other carnivores
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Decomposers
Get energy from detritus (no living organic material; leaves, wood, dead organism)
* include fungi and many prokaryotes * Important for recycling chemical elements
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Tropic structures
Structures of a community are determined by the feeding relationships between organisms
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Food chain
The transfer of food energy up the trophies levels
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Food ebs
Linked food chains
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Disrupted ecosystems
* if energy resources change so can the number and size of trophies levels (increase energy = increase tropic levels or decrease energy = decrease tropic levels) * A change at the producer level can affect the number and size of the remaining trophies levels
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Primary production
The amount of light energy that is converted to chemical energy
* set a spending limit for the entire ecosystems energy budget
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Gross primary production (GPP)
Total primary production in an ecosystem
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Net primary production (NPP)
the GPP minus the energy used by the primary producers for respiration (Ra)
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Secondary production
the amount of chemical energy in a consumer’s food that is converted to new biomass
* the transfer of energy between tropic levels is at 10% efficiency
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Biogeochemical cycles
Nutrient cycles that contain both biotic and abiotic factors
* water, carbon, nitrogen and phosphorus cycle
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Water cycle
Water is essential for all life and influences the rate of ecosystem processes
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Carbon cycle
Carbon is essential for life and required in the formation of organic compounds
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Nitrogen cycle
Nitrogen is important for the formation of amino acids, proteins and nucleic acids
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Phosphorus cycle
Phosphorus is important for the formation of nucleic acids, phospholipids and ATP (energy)
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Population
A group of individuals of the same species living in an area
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Population ecology
Analyzes the factors that affect population size and how and why it changes over time
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Density
The number of individuals per unit area
Can be determined by:
* counting number of indizaos in the population * Sampling techniques (count small areas, average the areas and then use the averages to estimate total population size)
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Dispersion
The pattern of spacing among individuals within a population
The study of the vital statistics of populations and how they change over time
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Life table
Age specific summary of the survival pattern of a population
* represented by a survivorship curve
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Type 1 curve
Low death rate during early/middle life and high death rate later in life
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Type II curve
Constant death rate over the lifespan of the organism
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Type III curve
High death rate early in life and lower death rate for those that survive early life
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Exponential growth model
A population living under ideal condition (ie easy access to food, abundant food, free to reproduce, ect)
* population grows rapidly * J shaped curve
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Logistic growth model
The per capita rate of increase approaches zero as the population size nears its carrying capacity
* the density of individual exceeds the system’s resource availability
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Life history
The traits that affect an organism’s schedule of reproduction and survival
Can be affect by:
* when reproduction begins * How often the organism can reproduce * The number of offspring produced per reproductive episode
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K-selection
* density dependent selection * Selection for life history traits that are se site to population density * Seen in high density populations that are close to carrying capacity (K)
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R-selection
* density independent selection * Selection for life history traits that maximize reproduction success * Seen in low density populations with little competition
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Density dependent regulation
As a population increases, factors can slow or stop growth by decreasing birth rate and increasing death rate