particular set of abiotic + biotic conditions and resources to which an organism or population responds
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fundamental niche
full range of conditions and resources in which a species could survive and reproduce
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realized niche
the actual conditions and resources in which a species exists due to biotic interactions
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biotic factors
living components of ecosystem -- organisms, interactions, waste -- that (in)directly affect another organism
1) every relationship that organism may have 2) where it lives 3) how it responds to resources available 4) how it alters these biotic factors
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abiotic factors
non-living physical factors that influence organisms and ecosystem -- eg temperature, sunlight, pH, salinity, pollutants
1) how much space there is 2) availability of light water, etc
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can two species inhabit the same ecological niche in the same space at the same time?
no; if many species live together, they must have slightly different needs and responses so they are not in the same niche
ex: lions and cheetahs both live in the African savannah, but hunt different prey -- lions take down zebras and buffalos, cheetahs take down gazelles and impalas
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limiting factors
prevent a community, population, or organism growing larger; will slow population growth as it approaches the carrying capacity of the system
ex: phosphate in limited supply in aquatic systems; low temp in tundra, freezing the soil and limiting water availability to plants
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population dynamics
the study of the factors that cause changes to population sizes
all interactions result in one species having an effect on the population dynamics of the others and on the carrying capacity of the others' environment`
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competition
two conditions applied jointly
1) all organisms in any ecosystem have some effect on every other organism in that ecosystem 2) any resource in any ecosystem exists only in limited supply
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intraspecific competition
between members of the same species
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what is true when numbers of a population are small?
there is little real competition between individuals for resources
provided numbers not too small for individuals to find mates, population growth will be high
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what happens as the population grows?
as the population grows, the competition between individuals for resources grows until carrying capacity of the ecosystem is reached
stronger individuals claim larger share of resources
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what are some strategies species employ when dealing with intraspecific competition?
being territorial, eg deer -- individual/pair holds area and fends off rivals
individuals most successful reproductively holds biggest territory, hence access to more resources, more successful at breeding
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what is the effect of intraspecific competition on population numbers?
intraspecific competition tends to stabilize population numbers
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how is intraspecific competition displayed on a graph?
it produces a logistic growth curve, which is s-shaped
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interspecific competition
individuals of different species could be competing for the same resource
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what could result from interspecific competition?
1) interspecific competition may result in a balance where both species share the resource 2) other outcome is that one species may totally out-compete the other (known as competitive exclusion)
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competitive exclusion
one species may totally out-compete the other
ex: garden overrun by weeds; number of weed species coexist together, but original domestic plants have been totally excluded
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predation
when one animal, the predator, eats another animal, the prey/consumption of one organism by another
ex: lions eating zebras, wolves eating moose
not only animals eat other animals -- some plants (insectivorous plants) consume insects and other small animals
a relationship between two species in which one species (the parasite) lives in or on another (the host), gaining food from it
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do parasites kill their host?
no -- not beneficial for them, but high parasite population densities can lead to the host's death
ex: vampires bats and intestinal worms
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mutualism
relation between two+ species in which all benefit and none suffer; example of symbiosis
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what are the types of symbiosis?
1) parasitism 2) commensalism: one partner is helped and the other is not significantly harmed 2ex) fern growing half-way up a tree trunk 3) mutualism
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what is a prime example of mutualism?
lichen: close association of fungus underneath and algae on top
fungus benefits by obtaining sugars from algae algae benefits from minerals + water fungus absorbs and passes on to the algae
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exponential/geometric growth
no limiting factors slowing growth
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s-curves
1) starts with exponential growth -- no limiting factors affect the growth at first 2) above a certain population size, growth rate slows down gradually, resulting in population of constant size
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carrying capacity
in s-curve, where numbers stabilize -- maximum number of load of individuals that an environment can carry/support
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environmental resistance
area between exponential growth curve and s-curve
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j-curve
see a boom and bust pattern 1) population grows exponentially 2) suddenly collapse (diebacks)
often, population exceeds carrying capacity on long-term or continuing basis before the collapse (overshoot)
note: long-term basis as carrying capacity can be exceeded in the short term (human race right now)
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what does the j-curve not show?
the gradual slow down of population growth with increasing population size
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what is j-curve typical of?
microbes, invertebrates, fish, small mammals
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community
group of populations living and interacting with each other in a common habitat
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ecosystem
community and physical environment it interacts with
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respiration
breaking down food, often in form of glucose, to release energy used in living processes
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what are the living processes?
MRS GREN
m - movement r - respiration s - sensitivity g - growth r - reproduction e - excretion n - nutrition
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what are the two types of respiration?
aerobic - oxygen anaerobic - no oxygen
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aerobic respiration
energy released and used waste products are carbon dioxide and water
c6h12o6 + 6o2 --> energy + 12h2o + 6co2
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what kind of energy is produced in respiration?
heat energy -- dissipated into the environment
increases entropy of the system while organism maintains relatively high level of organization (low entropy)
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photosynthesis
green plants convert light energy into chemical energy
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compensation point
rates of two processes equal (no net release of oxygen or co2)
neither adding biomass or using it up to stay alive -- maintaining itself
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food chain
flow of energy from one organism to the next
shows the feeding relationships between species in an ecosystem
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what do the arrows connecting the species indicate?
the direction of transfer of biomass and energy
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trophic levels
feeding levels of organisms
usually start with primary producer (plant) and end with carnivore at top of chain (a top carnivore)
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what are the two classifications of the way organisms obtain energy?
producers and consumers
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producers
1) autotrophs (green plants): make food from carbon dioxide and water using sunlight energy 2) chemosynthetic organisms: make food from simple compounds (ammonia, hydrogen sulphide, methane), do not require sunlight, often bacteria found in deep oceans
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consumers
also called heterotrophs
feed on autotrophs/heterotrophs to obtain energy (herbivores, carnivores, omnivores, detritivores, decomposers)
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what is the hierarchy of feeding?
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food chain
visual representations of feeding relationships between one organism and another
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ecological pyramids
graphical models of quantitative differences between amounts of living material stored at each trophic level of a food chain; usually measured for given area and time
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benefits of ecological pyramids
1) allow easy examination of energy transfers and losses 2) give idea of what feeds on what and what organisms exist at different trophic levels 3) help to demonstrate that ecosystems are systems in balance
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types of ecological pyramids
1) pyramids of numbers 2) pyramids of biomass 3) pyramids of productivity
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pyramid of numbers
number of organisms at each trophic level in food chain at one time (standing crop)
units: number/unit area
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what is the trend of pyramid of numbers?
length of each bar gives measure of relative numbers
most pyramids are broad at base and have many individuals in producer level, but some may have single large plant, a tree as producer, so base is one individual that supports many consumers
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advantages of pyramid of numbers
1) simple, easy method of giving an overview 2) good at comparing changes in population numbers with time or season
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disadvantages of pyramid of numbers
1) all organisms included regardless of size -- pyramid based on oak tree would be inverted 2) does not allow for juveniles or immature forms 3) numbers can be too great to represent accurately
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pyramid of biomass
contains biomass at each trophic level
units: mass per unit area (eg g*m^-2)
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biomass
quantity of dry, organic material in an organism, population, trophic level, or ecosystem
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what is the trend of the pyramid of biomass?
most likely to be a pyramid shape with some exceptions
ex: oceanic ecosystems where producers are phytoplankton (reproduce fast but present only in small amounts)
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advantage of pyramid of biomass
overcomes problems of pyramids of numbers
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disadvantages of pyramid of biomass
1) only uses samples from populations, so impossible to measure exactly 2) organisms must be killed to measure dry mass 3) time of year affects the result 4) energy content not represented
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what is true about both pyramids of numbers and biomass?
they are snapshots at one time and place, meaning that the pyramids vary with season and year
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pyramid of productivity
shows the rate of flow of energy or biomass through each trophic level
shows energy or biomass being generated and available as food to the next trophic level during a fixed period of time
units: energy or mass per unit area per period of time (J*m^-2*yr^-1)
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how is the pyramid of productivity different from the other two?
show the flow of energy OVER TIME
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what is always true about pyramids of productivity?
1) they are always pyramid-shaped in healthy ecosystems as they follow the second law of thermodynamics 2) each bar will be about 10% of the lower one (500 --> 50 --> 5)
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advantages of pyramids of productivity
1) most accurate system -- shows actual energy transferred and allows for rate of production 2) allows comparison of ecosystems based on relative energy flows 3) pyramids are not inverted 4) energy from solar radiation can be added
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disadvantages of pyramids of productivity
1) difficult and complex to collect energy data as rate of biomass production over time is required 2) still problem of assigning a species to a particular trophic level when they may be omnivores
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bioaccumulation
concentration of chemical high enough to cause disease/death
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biomagnification
when top trophic levels take in so much of chemical that causes disease or death
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which trophic level in a food chain is most susceptible to alterations in the environment?
top carnivores -- have limited diet, so change in prey has large effect
population numbers are low because fall in efficiency along a food chain, so ability to withstand negative influences is more limited than species lower in the food chain with larger populations
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trophic efficiency
10%
major part of energy used in respiration and lost as heat
less and less energy available after 4 trophic levels
this is why top carnivores are so vulnerable -- only so much energy available (too hard for them to accumulate enough energy to grow to large size and maintain bodies)
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solar constant
1400 watts per second
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what is the only way for life to convert solar energy into food?
photosynthesis by green plants
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productivity
conversion of energy into biomass over a given period of time
rate of growth or biomass increase in plants and animals
measured per unit area per unit time
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net productivity (NP)
gain in energy or biomass per unit area per unit time that remains after deductions due to respiration
results from fact all organisms have to respire to stay alive so some of the energy used in staying alive instead of being used to grow
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primary productivity
autotrophs base unit of all stored energy in any ecosystem
light energy converted into chemical energy by photosynthesis
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gross primary productivity (GPP)
total gain in energy or biomass per unit area per unit time by green plants -- energy fixed by green plants by photosynthesis
plants first organisms in production chain (fix light energy and convert to sugar) so possible to calculate plant's energy uptake by measuring amount of sugar produced
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why do we prefer NPP over GPP?
measuring sugar produced extremely difficult as much of it is used up by plants in respiration almost as soon as it is produced
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net primary productivity (NPP)
rate at which plants accumulate dry mass
measured in g*m^-2
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what happens to the glucose produced in photosynthesis?
1) provides for growth, maintenance, and reproduction with energy being lost as heat during respiration 2) remainder deposited in and around cells as new material represents stored dry mass -- store of energy is potential food for consumers within the ecosystem
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equation for NPP
NPP = GPP - R
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what is the theoretical max amount of energy available to all animals?
total amount of plant material
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what happens to the total amount of plant material?
1) lost from food chains as it dies and decays OR 2) eaten by herbivores, meaning it's removed from primary productivity
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how does the amount of biomass produced vary?
1) spatially: some biomes have higher NPP than others 1ex) tropical rainforest vs tundra 2) temporally: many plants have seasonal patterns of productivity linked to changing availability of basic resources (light, water, warmth)
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net secondary productivity (NSP)
total gain in energy or biomass per unit area per unit time by consumers after allowing for losses in respiration (also animals, but mainly respiration)
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what happens to energy that goes into the herbivore that doesn't make new biomass?
only food that crosses the gut wall of animals is absorbed and used to power life processes (assimilated food energy) a) some assimilated food energy used in cellular respiration to provide energy for life processes b) some removed as nitrogenous waste (urine) c) rest stored in dry mass of new body tissue
some ingested plant material will pass straight through the herbivore and be released as feces -- not absorbed and provides no energy
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equation for NSP
NSP = GSP - R
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gross secondary productivity (GSP)
total food ingested including food that turns into feces
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trophic level - carnivore
1) assimilate 80% of energy in diets 2) egest less than 20% 3) have to chase moving animals so higher energy intake offset by increasing respiration during hunting 4) biomass locked up in prey foods
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trophic level - herbivores
1) assimilate 40% of energy in diet 2) egest 60% 3) graze static plants
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flows of energy and matter
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major biogeochemical cycles (water, carbon, nitrogen, sulfur, phosphorous) characteristics
1) movement of matter, such as nutrients, through an ecosystem is very different from movement of energy 2) energy travels from the sun through food webs and eventually lost as heat 3) nutrients and matter are finite and recycled and reused (via decomposer food chain) 4) organisms die and are decomposed and nutrients are released, eventually becoming parts of living things again, when taken up by plants
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where is the carbon stored?
in carbon or carbon dioxide sinks
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carbon (dioxide sinks) may be:
1) organic (complex carbon molecules 1a) organisms in the biosphere (plants and animals) 1b) fossilized life forms (fossil fuels)
2) inorganic (simple carbon molecules) 2a) locked up or fixed into solid forms and stored as sedimentary rocks and fossil fuels (stored and locked up for millions of years) 2b) oceans where carbon dissolved or locked up as carbonates in shells of marine organisms 2c) soil 2d) small proportion is carbon dioxide in the atmosphere
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carbon cycle
carbon circulates through living and non-living systems in the ecosphere