Specialized in taxonomy, developed a binomial system of naming organisms according to genus and species that is still used today. Also adopted a system for grouping similar species into a hierarchy of increasingly general categories.
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James Hutton (1726-1797)
Gradualism, the idea that earth is continuously, slowly modified by the accumulation of many small geological changes such as erosion, wind, sedimentation, etc.
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Uniformatarianism, the theory that changes in the earth's crust during geological history have resulted from the action of continuous and uniform processes.
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Comte de Buffon (1707-1788)
Recognized the dynamic relationship between the external environment and living forms. Also recognized vestigial structures.
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Jean-Baptiste Lamarck (1744-1829)
The first modern European thinker to develop a comprehensive theory of evolutionary change.
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Idea that characteristics are acquired during an individual organism's lifetime and could be inherited
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Georges Cuvier (1769-1832)
Founded comparative anatomy and paleontology; demonstrated the fact of organic extinction.
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Realized that extinction was a common occurence, advocated catastrophism even though he was against evolution.
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Alfred Russel Wallace (1823-1913)
English naturalist who proposed, independently of Charles Darwin, the concept of natural selection as a mechanism for evolution and as a way to explain the great variety of living things
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Charles Darwin (1809-1882)
Theory of natural selection, a plausible mechanism for biological evolution
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Argued that all the organisms that have ever lived arose through descent with modification, the evolutionary alteration and diversification of ancestral species
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Gregor Mendel (1822-1884)
Investigated inheritance in pea plants by observing differences in various traits of the plants from generation to generation, leading to the idea that information needed to build an organism is packaged into units associated with specific traits
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Gradualism
The view that Earth and its living systems changed slowly over its history
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Essentialism
a belief that things have a set of characteristics that make them what they are, and that the task of science and philosophy is their discovery and expression; the doctrine that essence is prior to existence.
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Scala Naturae
all living organisms arranged in a linear order from simple to complex
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Catastrophism
The theory that Earth has been affected by sudden, violent events that were sometimes worldwide in scope
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natural selection
The evolutionary process by which alleles that increase the likelihood of survival and the reproductive output of the individuals that carry them become more common in subsequent generations.
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Homology
Similarity in characteristics resulting from a shared ancestry.
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homologous structures
Characteristics that are similar in two species because they inherited the genetic basis of the trait from their common ancestor
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transitional forms
fossils that connect ancestral species with their descendants through a series of tiny steps
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Archeopteryx
Famous bird-like fossil that links birds and reptiles
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Tiktaalik
"missing link" thought to be a transitional form between lobefin fish and tetrapods
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Pakicetus
52 million year old whale ancestor
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Dorudon
An extinct species that had a vestigial pelvis and is related to whales
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law of succession
fossil species are strikingly similar to living species in the same geographic areas
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genetic structure
frequency of different alleles and genotypes in a population
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gene pool
The sum of all alleles at all gene loci in all individuals in a population
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gene locus
The particular site on a chromosome at which a gene is located.
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phenotypic frequency
The percentage of individuals in a population possessing a particular phenotype
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genotypic frequency
The percentage of individuals in a population possessing a particular genotype
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allele frequency
The abundance of one allele relative to others at the same gene locus in individuals of a population.
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Microevolution
Small-scale genetic changes within populations, often in response to shifting environmental circumstances or chance events.
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Macroevolution
Large-scale evolutionary patterns in the history of life, producing major changes in species and higher taxonomic groups
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Hardy-Weinberg Principle
An evolutionary rule of thumb that specifies the conditions under which a population of diploid organisms achieves genetic equilibrium
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Hardy-Weinberg equilibrium
condition that occurs when the frequency of alleles in a particular gene pool remain constant over time
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genetic drift
Random fluctuations in allele frequencies as a result of chance events; usually reduces genetic variation in a population.
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bottleneck effect
An evolutionary event that occurs when a stressful factor reduces population size greatly and eliminates some alleles from a population.
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founder effect
An evolutionary phenomenon in which a population that was established by just a few colonizing individuals has only a fraction of the genetic diversity seen in the population from which it was derived.
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gene flow
The transfer of genes from one population to another through the movement of individuals or their gametes.
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nonrandom mating
changes in a population's gene pool due to mate choice or other forces that cause individuals to mate with certain phenotypes more than others
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Inbreeding
A special form of nonrandom mating in which genetically related individuals mate with each other.
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assortive mating
Mating of individuals with similar phenotypes
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Polymorphism
The existence of discrete variants of a character among individuals in a population.
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morph
The form or shape of an organism, or of a part of an organism.
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gel electrophoresis
Technique by which DNA, RNA, or protein molecules are separated in an agarose gel subjected to an electric field.
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geographical variation
differences in the phenotypic variation between populations that are separated geographically
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cline
A pattern of smooth variation in a characteristic along a geographical gradient.
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balanced polymorphism
The maintenance of two or more phenotypes in fairly stable proportions over many generations.
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frequency-dependent selection
A form of natural selection in which rare phenotypes have a selective advantage simply because they are rare.
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Darwininan Fitness
Relative contribution an individual makes to the gene pool of the next generation compared to contribution by other individuals
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relative fitness
The number of surviving offspring that an individual produces compared with the number left by others in the population.
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stabilizing selection
A type of natural selection in which individuals expressing intermediate phenotypes have the highest relative fitness.
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directional selection
A type of natural selection in which individuals near one end of the phenotypic spectrum have the highest relative fitness.
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diversifying (disruptive) selection
A type of natural selection in which extreme phenotypes have higher relative fitness than intermediate phenotypes
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sexual selection
A form of natural selection established by male competition for access to females and by the females' choice of mates.
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sexual dimorphism
Differences in the size or appearance of males and females.
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Macroevolution
Large-scale evolutionary patterns in the history of life, producing major changes in species and higher taxonomic groups over long periods of time
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Anagenesis
The slow accumulation of evolutionary changes in a lineage that continues to exist as an interbreeding population
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Cladogenesis
Branching evolution that occurs when one or more new species branches out from a parent species
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morphological species concept
The concept that all individuals of a species share measurable traits that distinguish them from individuals of other species
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biological species concept
The definition of species based on the ability of populations to interbreed and produce fertile offspring
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Ernst Mayr
Fused Charles Darwin's theory of natural selection with the modern science of genetics; coined the "biological species" concept, altering understanding of the nature of a species
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geographic isolation
physical separation of a group of individuals from others of the same species
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reproductive isolation
Separation of species or populations so that they cannot interbreed and produce fertile offspring
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prezygotic barriers
A reproductive isolating mechanism that acts before the production of a zygote, or fertilized egg
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habitat (ecological) isolation
A prezygotic reproductive isolating mechanism in which species that live in the same geographical region occupy different habitats
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behavioral isolation
A prezygotic reproductive isolating mechanism in which two species do not mate because of differences in courtship behavior; also known as ethological isolation.
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temporal isolation
A prezygotic reproductive isolating mechanism in which species live in the same habitat but breed at different times of day or different times of year.
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mechanical isolation
A prezygotic reproductive isolating mechanism caused by differences in the structure of reproductive organs or other body parts.
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gametic isolation
A prezygotic reproductive isolating mechanism caused by incompatibility between the sperm of one species and the eggs of another; may prevent fertilization.
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postzygotic barriers
A reproductive isolating mechanism that acts after zygote formation
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reduced hybrid viability
A postzygotic reproductive isolating mechanism in which a hybrid individual has a low probability of survival to reproductive age
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reduced hybrid fertility
A postzygotic reproductive isolating mechanism in which hybrid offspring cannot form functional gametes
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hybrid breakdown
A postzygotic reproductive isolating mechanism in which hybrids are capable of reproducing, but their offspring have either reduced fertility or reduced viability
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Reinforcement
The enhancement of reproductive isolation that had begun to develop while populations were geographically separated
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allopatric speciation
The evolution of reproductive isolating mechanisms between two populations that are geographically separated.
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sympatric speciation
Speciation that occurs without the geographical isolation of populations.
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peripheral isolates
relatively small isolated or semi-isolated populations distributed on the periphery of a larger, central population; conditions favor allopatric speciation
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adaptive radiation
A cluster of closely related species from a common ancestor that are each adaptively specialized to a specific habitat, food source, or ecological niche; very common in "island-like" situations
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allopolyploidy
The genetic condition of having two or more complete sets of chromosomes from different parent species; may occur when two closely related species hybridize and produce polyploid offspring (may be initially sterile)
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Autopolyploidy
The genetic condition of having more than two sets of chromosomes from the same parent species; may occur when gametes spontaneously receive the same number of chromosomes as a somatic cell during meiosis
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parapatric speciation
when a species is spread out over a large geographic area but only reproduces with local species, resulting in the development of a new species
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Phylogeny
Evolutionary history of a species or group of species.
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Clade
A monophyletic group of organisms that share homologous features derived from a common ancestor.
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monophyletic
a grouping consisting of a clade that includes a most recent common ancestor and ALL of its descendents
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paraphyletic
grouping consisting of a clade that includes a most recent common ancestor and most but not all of its descendants
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polyphyletic
a grouping consisting of species that does not include their most recent common ancestor and also not including all the descendants of that ancestor
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Homology
Similarity due to descent from common ancestor; may serve different function in descendants
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Analogy
Similarity due to same function/evolutionary need
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convergent evolution
Process by which unrelated organisms independently evolve similarities when adapting to similar environments
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Synapomorphy
shared derived character; evidence of recent relationship
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Symplesiomorphy
shared ancestral character; evidence of more distant evolutionary relationship
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character
A heritable feature that varies among individuals
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character state
One of two or more distinguishable variants of a character
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ingroup
a group of closely related taxa that is investigated for evolutionary relationships
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outgroup
a reference group or a taxon outside the group of interest and more distantly related to the ingroup
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parsimony
A particular trait is less likely to evolve independently in separate evolutionary lineages rather than as a result of shared ancestry; hence, all other things being equal, the best hypothesis is the one that requires the fewest evolutionary changes
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Reducing atmosphere
A popular view about the early atmosphere that suggests that the atmosphere was mainly composed of CH4, NH3, hydrogen, water vapor and very little oxygen. These chemicals act as reducing agents where electrons are donated and allow chemical synthesis (electron bonding).
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Stanley Miller
An American Scientist who experimented in Harold Urey's laboratory at the University of Chicago in 1953 to see if the gases hypothesized to exist on early earth could really create organic compounds. Simulated the water cycle on early earth, isolating amino acids and nitrogenous bases in the process which can form polymers under certain conditions
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Fossils
The preserved remains or traces of organisms that once lived on Earth.