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What are the 3 domains of life
bacteria, eukarya, archae
serial endosymbiosis
how eukaryotes evolved from prokaryotes
endosymbiosis
A process where one cell(endosymbiont) lives within another cell(the host)
How did mitochondria evolve
aerobic bacteria was swallowed by a more complex archaea
How did chloroplats evolve
photosythetic bacterium engulfed by ansetral prokaryote
Proof of serial endosymbiosis
1. circular genome chromosomes like bacteria
Replicate simialr to bacteria
organelles transcribe their own DNA
Inner membranes have enzymes and transport systems similar to bacteria
ribosomes similar to bacteria
Why study plants
oxygen, food, beverage, animal feed, drugs, fibers
Linnean hierarchy
narrow to broad; species, genius,family, order, class, phylum, kingdom
Genera naming
genus+aceae
Species
Group of individuals that when they interbreed:
➢ produce fertile progeny without artificial
intervention
Subspecies
Capable of interbreeding but not in nature human intervention neededp of individuals
Intergenetic hybrids
fertile progeny involving 2 genus, X is used to indicate genus cross
plant histoyr
non vascular plants (mosses), vascular plants(ferns/seedlings), Vascular plants(flowers
angiosperms consist of 3 thinsg
flower, fruit, double fertilization
Monocot
1 cotyledon, flower parts in 3s, paralle vines, primary root replaved by adventious root, scatterd vasculature, 1 pollen pore, corn
Dicot eudicot
2 cotyledon, 5-4 flower parts, network of viens, ring formation vasculature, primary root/tap root, 3+ pollen pore
Alternating generamition
alternating between sporophyte, and gametophyte
mitosis
one cell divides, and produces 2 identical daughter cells diplpoid cell (2 chromosomes)
meiosis
one cell divides and produces 4 cells with ½ number of chromosomes (haploid) spores
sporophyte produces
spores haploid
gametophyte
produces gametes haploid
Alternation generation
uses both mitosis and meiosis alternate between the sporophyte and gametophyte generation
Male and female gametophytes in gymnosperms(non flowering) what is the sporophyte
pollen cones, ovulate cones, main body of plant with stems
Male and female gametophytes in angioosperms(flowering) what is the sporophyte
ouvle, pollen, main body of plant with stems
What are seed plants dependant on
sporophyte tissue for nutrition
In mosses what generation is predominant
gametophyte
In ferns what generation is predominant
50/50
In gymnosperms and angiosperms which is more predominant
sporophyte
What is the main life cycle in angiosperms
Adult (sporophyte) phase is the main life
cycle phase
Angiosperms are heterosporous
they produced 2 spores, microspores (male→pollen grains), megaspores(female→ ovule)
Stamen male parts
anther and filament
carpels female parts
stigma, style, ovary
Angiosperm life cycle
In anthers microsporocyte divide me to produce 4(n) microspores
In ovule megasporocyte divide me produce 4(n) megaspores ¼ survives
microspores divide mit produce pollen grain
megaspores divide mit, 7(n) eggs, synergids, antipodals, polar nuclei
pollen grain lands on stigma and pollinates
Pollen grain germinates tube cell creates tube down style, generative cell follows 2 sperms in generative cell, 1 fertilizes egg (2n zygote) other fert polar nuclei(triploid 3n)
zygote turns into embryo, triploid turns into endosperms
seed germinates, sporophyte matures cycle repeats
A flower is a reproductive structure that
produces gametes (ovule, pollen, egg & sperm)
➢ protects and nourishes the
megaspore gamete (carpel)
➢ captures the male gametes
(pollen)
➢ nourishes the embryos
➢ develops seed(s) and fruit(s)
What is a seed
An embryo and a nutrient source
surrounded by a protective coat
What is a fruit
enlarged ovary that contains seeds
The flower is the ———— shoot
of an angiosperm
reproductive
A complete flower consists of 4 whorls
sepals mod leaf
petals mod leaf
stamens
carpels
Incomplete flower
lack 1+ whorl
Inflorescence
cluster of flowers, wild carrot
Perfect flowers
flowers that contain both male and female parts
Incomplete flowers:Monoecious
male and female flowers are separate but on the same plant (zuchinni)
Incomplete flowers: Dioecious plants
male and female flowers are found on separate plants (willows)
What do pollen grain walls contain
sporopollenin, very hard material make for good fossils
Pollination vectors and percentages
Animals 80% Abiotic 20%-wind 98%-water2%
how many plants are animal pollinated
250,000 and pollinated by 100,000 animals including invertebrates and vertebrates
Flower methods of attraction
flower colour
nectar
odour
deception/mimicry
timing of pollen
flower shape
*may be more metabolically efficient than relying on wind→ creating lots of pollen)
Nectar and pollen rewards
pollen protein, nectar sugar 13 amino acids
Characteristics of flowers pollinated by birds
-range from scarlet to red to orange and generally lack nectar guides
➢ have very deep tubes usually without a landing platform, are pendant or horizontal, and
➢ have abundant nectar but emit no odor,
Bats as pollinated flower characteristics
-open at night, similar to moth-pollinated flowers
➢ are positioned below the foliage of the plant
o hanging pendant on a long pedicel or attached to the trunk or low limbs
➢ are drab white, green, or purple, bats are color-blind
➢ exude a strong musty odor at night, fermenting fluid, cabbage, or bats themselves
➢ large and tough, with lots of pollen and nectar
some have more than 1300 anthers and 7 to 15 ml nectar
Co-evolution
plants and the animals that pollinate them have co-
evolved
• in some cases, the flowers have shapes that are only accessible to certain pollinators buff-tailed sickle bills
Wind pollinated flowers tend to be
small, colorless, odorless, and lacking in nectar, lacking petals or have petals reduced to small scales, have flowers dangle in or wave
Where do grasses and sedges position their flowers
above leaves so they are exposed to the wind
Why are wind-pollinated pollen grains (frisbee shaped) smaller than animal-pollinated pollen grains
so they can be carried through the wind more easily 20-60 micrometers compared to 13-300
where are the flowers in corn
tassel staminate flowers, ear female flower, silk is stigma
Self-pollination/Selfing (10-13% of plants)
transfer of pollen from the anther to the stigma in the same flower and transfer of pollen from one flower to another on the same plant
Cross-Pollination or outcrossing
involves transfer of pollen from one genetically distinct plant to the stigma of another, increases the genetic diversity in the offspring
Selfing advantages
widespread propagation of the same, adapted, genotype
➢ no need for wind or pollinators
➢ little “metabolic cost”
o little wasted pollen
o no cost to produce nectar or other characters to attract pollinators
3 traits of angiosperms
Flowers
Fruits
Double fertilization
Disadvantages of self-fertilization
➢ restricted opportunity to create new genetic
combinations
-limited genetic diversity might lead to local extinction if pest & growing conditions change
• inbreeding depression
• loss of vigour
➢ may be caused by expression of deleterious
traits determined by recessive alleles
Homozygous individuals-AA or aa
Mechanisms that promote cross pollination
Genetic incompatibility
the ability of a plant to reject its own pollen and sometimes the pollen of closely related individuals based off of
→ Gametophytic self-incompatibility: In gametophytic self-incompatibility, the S-allele in the pollen genome governs the blocking of fertilization s1 cant fertilize s1s2 flower pollen tube will not germinate
→Sporophytic SI: fertilization is blocked by S-allele gene products in tissues of the parental sporophyte that adhere to the pollen grain wall, due to the parental tissue attached to the pollen wall.
A seed consists of and is
An embryo and nutrients surrounded by a protective coat, mature ovules that contain the
embryonic plants of the next generation
Oldest seed germination is
32,000 recovered from Siberian permafrost, silene stenophylla
Seeds and fruits are filled with stored foods intended to
→help the embryo germinate and grow
➢ or to attract an animal to eat the fruit and inadvertently carry seeds away
What vegetables are also considered fruits
string beans, eggplant, okra, tomato, squash, cucumber
Simple fruits
develop from a single or several fused carpels of one flower pea, fleshy or dry, can be dehiscent (open or discharge seeds) or indehiscent (they don’t open or discharge seeds) The thin skin derived from the outer layer of the ovary. The soft
fleshy skin is derived middle layer of the ovary and stony pit is derived from the inner tissue layer of the ovary.
Aggregate fruits (single flower multiple carpels(ovaries)
This process starts with a flower that contains several carpels with an equal number of ovaries. Each ovary contains a single ovule that develops into a seed following fertilization. As the ovaries develop and increase in size they attach. Each fruit in the aggregate is a drupe containing a stony pit.
Multiple fruits (develop from inflorescence)
ovary walls thicken they fuse together and become incorporated into 1 fruit
Accessory fruit (develops from tissue other than ovary)
strawberry fruit is enlarged receptacle tissue
➢ the dry “seeds” embedded in the
red tissue are achenes (or true
fruits) similarly, an apple is made up of
mostly enlarged hypanthium tissue
(which connects sepal, petals and
stamens in the flower)
➢ the core of the apple is the fruit
Fruits that develop without fertilization are ______ and called_____
seedless, parthenocarpic grapes, eggplant, watermelon
Plants that do not require pollination or other stimulation to produce parthenocarpic fruit exhibit
vegetative parthenocarpy
some plants, pollination (with dead pollen) or another stimulation is required for parthenocarpy called
Stimulative parthenocarpy
Seed wind dispersal characteristics
winged, plumed, ballistic dispersal
Animal seed dispersal techniques
attachment strategies, food reward
What is scarifying
breaking seed coat to speed up germination
The only stored food for seedling is
the endosperm and cotyledons
it costs the plant more to make seeds and fruits than it does to make
vegetative organs
Impermeable seed coats are broken by
passage through the animal tract, mechanical nicking freezing, thawing
Steps in germination eudicots
Water intake through micropyle
Enzymes break down food stored in cotyledon, which are transported to the radical for growth, radical pushes out
a hook forms in the hypocotyl, and growth pushes the hook above ground
the hypocotyl and radicle start growing and emerge out of the seed
the brown seed coat starts to peel off from top to bottom
hypocotyl from the seed appears green and is bent downward extends to the radicle that starts to develop into a root
The cotyledons are pushed half above the soil the hypocotyl straightens, pulling the cotyledons completely out
• foliage leaves emerge from the cotyledons and light straightens the shoot tip
Germination is the apperance of the
radical and shoot from the seed
Emergence is the apperance of the
seedling shoot above the soild surface
epigeal
cotyledons emerge from the ground
hypogeal
cotyledons remain in soil
Monocot seed germination
coleoptile pushes up through the soil creating a tunnel for the shoot tip to grow through
coleoptile and radicle start growing
and emerge out of the seed
The coleoptile faces upward and
radicle faces downward
the radicle develops into a root
below the soil
the shoot grows straight up through the tube of the coleoptile to establish the foliage leaves of the
seedling
Vascular plants have 3 organs
leaves, stems, roots
Root and shoot dependency
shoots rely on water and minerals absorbed by the root system and roots rely on sugar produced by photosynthesis in the shoot system
Plant cell features
membrane-bound nucleus and organelles like edoplasmic reticulum, golgi, mitochondria, but are unique and have cell wall, chloroplats, and central vacuole
The microfibrils are embedded within a noncellulose matrix, which can account for 60 -80% of primary cell walls, composed of
hemicelluloses (branched polysacchride) pectins(hetergenous mixture of sugars rich is galacturonic acid), extensins(glycoproteins)
microfibrils
celluslose in orthogonal layer
What is chlorophyl
chlorophyl is the green pigment in chloroplast which captures light
Where do light harvesting reactions take place
thylakoid membrane
Where does sugar synthesis take place
in stroma
Plant tissue falls into 2 categories
Meristematic and permanent tissue.
Meristematic tissue cells are either undifferentiated or incompletely differentiated; they continue to divide and contribute to the growth of the plant.
Permanent tissue consists of plant cells that are no longer actively dividing.
What are meristems
plant regions where there is cell division and continuous growth
What are the 3 meristematic tissues
apical, intercalary, lateral meristems
Apical meristems found and do what
located at the tips of stems and roots, which enable a plant to extend in length
Lateral meristem found and do what
found in the vascular cambium of stems facilitate growth in thickness or girth in a maturing plant
Intercalary meristems found and do what (monocots only)
occur at the bases of leaf blades and at nodes enable monocot leaf blades to increase in length from the leaf base i,e lawn grass
Primary meristems 3 permanet tissue types
dermal(covers protects plant), ground (site for
photosynthesis, provides a supporting matrix for
the vascular tissue, store water
and sugars), vascular(transports water, minerals, suagrs)